PALYNOLOGICAL ZONATION OF MID-PALAEOZOIC, NW SPAIN 



155 



but in the original description Rodriguez separated Amicosporiies 

 infraomator from A. subornata on the distal position of the dark 

 area. Cramer & Diez ( 1 975 ) on the other hand thought that the dark 

 ring in A. jM/70/-nfl/r)r was proximal. If further specimens show that in 

 A. siibornatorthe dark ring is distal then the two species may have to 

 be combined. 



Remarks. The spores differ from typical members of the genus 

 Amicospohtes in having a proximal kyrtome (see under C. cromatica). 

 Rodriguez ( 1978b) included in her species both spores with a distal 

 annulus, not well defined at the inner margin, and others where the 

 annulus was dissected into a series of verrucate-like structures 

 (Rodriguez, 1978b: pi. 1, fig. 12). 



Occurrence. Geras, upper RS, H and lower EC Biozones, La 

 Vid, upper RS to top H Biozones, Argovejo, RS, H, EC and MN 

 Biozones, La Peral upper H and lower EC Biozones; Pfidoli and 

 lowermost Devonian. 



Infraturma PATINATI Butterworth & Williams 1958 emend. 



Smith & Butterworth 1967 



Genus CHELINOSPORA Allen 1965 



Type species. Chelinospora concinna Allen 1965. 



Chelinospora canistrata sp. nov. PI. 8, fig. 8. 



Derivation of name. Latin canistmm n. wicker basket, canistrata 

 resembling a basket. 



HOLOTYPE. FM 1492, sample Geras 92/5, slide 2 (542), co-ord. 

 083 1024, E.F. no. J33/1 ; Lower San Pedro Formation, Geras section. 

 Province of Leon, Cantabrian Mountains, northwest Spain. 



Diagnosis. A Chelinospora with low large verrucae-muri and 

 closely packed convolute microrugulae. 



Description. Proximally hilate, distally patinate miospores; hilum 

 thin, sometimes diaphanous, frequently torn, or may be collapsed or 

 lost, with a narrow concentric fold just inside the crassitude; narrow 

 equatorial curvatural crassitude with short proximal radial muri 

 extending to the hilum margin seen on a few specimens; distal 

 sculpture of low verrucae and muri, often indistinct, and closely 

 packed sinuous microrugulae < 1 pm wide. Y-mark with narrow 

 'folds' extend c. 2/3 spore radius and merge into crassitude at the 

 hilum margin. 



Dimensions. 25-43 ^m (based on 18 specimens). 



Comparisons. The sculpture of low verrucae-muri and 

 microrugulae distinguish Chelinospora canistrata sp. nov. from 



other Chelinospora species. 



Occurrence. Found in all sections apart from La Vid, uppermost 

 RS, H, and lower EC Biozones. 



Chelinospora cantabrica sp. nov. PI. 4, fig. 7; PI. 9, fig. 1 



Derivation of name. Named after the Cantabrian region, NW 

 Spain. 



HOLOTYPE. FM 1493 (PI. 9, fig. 1), diameter 39 |jm, sample 

 Argovejo 92/5a, slide 3 (473), co-ord. 1441082, E.F no. P39/1; 

 Lower San Pedro Formafion, Argovejo section. Province of Leon, 

 Cantabrian Mountains, northwest Spain. 



Diagnosis. A Chelinospora with a reticulum composed of broad 

 muri with occasional constrictions and narrow sutures, lumina wide, 

 polygonal to irregular. 



Description. Proximally hilate miospores, equatorially 2-A \xm 

 thick, distal exine thick (not measured); amb subcircular to 

 subtriangular. Proximal exine thin, frequently broken, or absent, 

 trilete mark not seen. Equatorially and distally sculptured with a 

 broad reticulum, muri 1-10 pm wide, 1-3 urn high, lumina wide and 

 polygonal 2.5-10 |im wide; muri show constrictions broken by 

 narrow sutures at the junction with adjacent polygonal fields. 



Dimensions. 5-51 |am (based on 20 specimens). 



Comparisons. The broad muri with distinct constrictions and 

 sutures, and wide +/- polygonal lumina, distinguish this species 

 from other species of Chelinospora. Coronaspora reticulata sp. nov. 

 has similar reticulate sculpture but has an equatorial crassitude and a 

 proximal kyrtome. 



Occurrence. Argovejo, Geras, and La Vid sections, Lower San 

 Pedro Formation, RS and lowermost H Biozones, occasional finds in 

 higher zones in the La Vid section. 



Chelinospora cassicula Richardson & Lister 1969 



1969 Chelinospora cassicula Richardson & Lister: 242-43, 



pi. 42, figs 10-12. 

 1978a Chelinospora mariae Rodriguez (pars); Rodriguez: 10. pi. 



1, fig. 3 only. 



Remarks. Rodriguez, 1 978a (pi. 1 , figs 3-5 ) placed several spore 

 morphologies in this taxon but from the illustration the holotype 

 belongs in C. cassicula. 



In the Spanish material examined for this paper several 

 Chelinospora species with an equatorial band of radial muri form an 

 evolving 'lineage" (morphon) beginning with C hemiesferica. 



PLATE 11 



Fig. 1 Conochitina pachycephala Eisenack 1964. EC 901, lateral view, x 170, LP3/SB4/45. 



Fig. 2 Conochitina alargada Cramer 1 967. EC 9 1 , lateral view, x 220, Arg4/SB7/3. 



Figs 3, 5, 6 Cyathochitina elenitae Cramer 1964. 3, EC 92, lateral view, x 200, LP3/SB4/48; 5, EC 93. lateral view x 200. Arg4/SB7/3; 6, EC 94. Arg3/ 



SB7/1; 6a, lateral view x 150; 6b, detail of carina, x 750. 

 Fig. 4 Spinachitina sp. EC 95, LP2/SB4/43; 4a, lateral view, x 300; 4b, basal margin detail, x 1 750. 



Calpichitina velata (Wrona 1980). EC 96, Gerl5b/SBGeras-l; 7a, apertural view, x 500; 7b, detail of operculum x 1000. 

 Vinnalochilina horrenlis (Jaglin 1986). EC 97, Gerl3/SBGeras-2: 8a, apertural view, x 500; 8b, detail of operculum x 1000. 

 Cingiilochitina wronai Paris & Kri'z 1984. EC 98. lateral view, x 500, Arg 7/54 1/2, (R/34). 

 Umochitina imm Eisenack 1934. EC 99. lateral view, x 300. LP20/SB5/55. 



Cingulochitina ervensis (Paris 1979). EC 100, Gerl 1/458/7, (N31/2): 11a. lateral view, x 250; lib, detail of carina, x 1000. 

 Cyathochitina sp. B, Paris 1981. EC 101, Arg3/SB7/1; 12a, lateral view, x 300; 12b. detail of carina, x 800. 

 IMargachitina catenaria Obut 1973. EC 102, apertural view, x 400, Gerl6/471/3. 

 Eisenackitina bohemica (Eisenack 1934). EC 103. lateral view, Gerl4/468/14. (Q26/4). 



Fig. 7 

 Fig. 8 

 Fig. 9 

 Fig. 10 

 Fig. 11 

 Fig. 12 

 Fig. 13 

 Fig. 14 



