156 



J.B. RICHARDSON, R.M. RODRIGUEZ AND S.J.E. SUTHERLAND 



followed by C. cf. hemiesferica in the H Biozone, accompanied by 

 C. lavidensis in the upper part of the H zone. Chelinospom media 

 and C. cassicula are confined to the higher parts of the section 

 studied Lower Lochkovian (upper EC & lower MN Biozones). 



Occurrence. Argovejo, Geras, and La Vid sections Upper San 

 Pedro Formation, lower MN (NA) Sub-Biozone. 



Chelinospora hemiesferica (Cramer & Diez) comb. nov. 



PI. 4, fig. 9; PI. 9, figs 3. 5 



Basionym. Iberoespora hemiesferica Cramer & Diez, 1975: 341 , 

 pi. 2, figs 34-36. 



1 978c Cymbosporites dittonensis Richardson & Lister; Rodriguez: 

 pi. 3, fig. 18. 



HOLOTYPE AND LOCUS TYPICUS. Cramer & Diez 1975: 334, 336; 

 pi. 2, figs 34, 35: 'base of a thick green shale 10 m below the exposed 

 top of the San Pedro Formation at the westernmost outcrop in the 

 village of Argovejo in the province of Leon, Spain'. 



Emended diagnosis. A Chelinospora with closely spaced nar- 

 row muri, geniculate in plan, over the distal hemisphere and an 

 equatorial to subequatorial area, where the muri become radial. 

 Proximal exine thin, trilete mark labrate, labra narrow. 



DIMENSIONS. 30^0|jm (Cramer & Diez, 1975): 29^4 |jm (based 

 on 75 specimens, present study), muri <1 \xm wide and high. 



Remarks. The spores may be double-layered and on some speci- 

 mens a membranous curvatural zone is seen bearing radial extensions 

 of the distal muri. 



Comparison. The tightly packed geniculate narrow distal muri 

 distinguish this species. Cramer & Diez (1975) refer to a 'pro- 

 nounced sculpture of brain-like muri' distally. 



Occurence. Distributed through the H to lower MN Biozone 

 (NA Sub-Biozone). 



Chelinospora cf. hemiesferica (Cramer & Diez) comb, 

 nov. PI. 4, figs 6, 10; PI. 9, fig. 4; PI. 10, fig. 1 



Description and comparison. As for C. hemiesferica, but with 

 broader, more widely spaced distal muri; muri more than 1-3 |jm 

 wide and 1-2 |am apart. 



Occurrence. Found in all sections, Argovejo, Geras, La Peral 



and La Vid sections. Upper San Pedro Formation; range hemiesferica, 

 EC Biozones to lower MN (NA) Sub-Biozone; found in the 

 hemiesferica Biozone only in the La Peral Section. 



Chelinospora lavidensis sp. nov. 



PI. 9, fig. 2 



1978c Iberoespora hemiesferica Cramer & Diez; Rodriguez: 4 1 8, 



pi. 3, fig. 15. 

 1983 Dicryotriletes geriense Rodriguez: pi. 6, fig. 6 only. 



Derivation of name. Named for the La Vid locality. 



HOLOTYPE. FM 1494(P1.9, fig. 2), diameter 33 |jm; sample Ger 9, 

 sHde Dl/1, co-ord. 195 1056; E.F. no. U/36, upper San Pedro 

 Formation, Geras section. Province of Leon, Cantabrian Mountains, 

 northwest Spain. 



Diagnosis. A Chelinospora with narrow low muri, 1 jim or less 

 wide and >l|im high, forming an irregular reticulum with broad 

 lumina. 



Description. Proximally hilate miospores, equatorially 2-A pm 

 thick, distal exine thick (not measured), amb subcircular to sub- 

 triangular. Proximal hilum thin, frequently broken or absent, trilete 

 mark not seen. Equatorially and distally sculptured with an irregular 

 reticulum, muri 1 ^im or less wide, <1 pm high, lumina highly 

 variable in shape and size on a single spore, at the distal pole large 

 and polygonal 8-10 pm wide, subequatorially c. 1-2 \im wide; muri 

 become radial at the equator. Y-mark with narrow 'folds' extends c. 

 2/3 spore radius and merge into crassitude at the hilum margin, 

 frequently gape over much of the hilum forming a distinct triangular 

 area. 



Dimensions. 29^3 pm (based on 17 specimens). 



Comparison. Chelinospora hemiesferica (Cramer & Diez) comb, 

 nov. has a distal sculpture of closely packed geniculate muri but 

 resembles C lavidensis sp. nov. because both species have an 

 equatorial band of radial muri. 



Remarks. Several species of Chelinospora (C. hemiesferica, C 

 sanpetrensis. C. media and C. lavidensis) have a subequatorial band 

 of muri normal to the equator, which may indicate affinity or 

 possibly represents a feature related to the development of the tetrad 

 in murornate spores. 



Occurrence. Upper part of H and EC Biozones, rare in lower 

 MN (NA) Sub-Biozone, Upper Pfidoli and lowermost Devonian, 

 Argovejo, Geras and La Vid sections. 



1 A/508/12. (Q27/4); 2, PC 105. LP20/SB5/54; 2a, lateral 



PLATE 12 



Figs 1, 2 Pseudoclathrochitina carmenchui (Cramer 1964). 1, PC 104. lateral view, x 350. Argl 



view X 400; 2b, basal margin detail, x 1 100. 

 Fig. 3 Margachitina elegans var. carta (Cramer 1964). PC 106, lateral view, x 300, Argl2/5I0/8, (K30/4). 

 Fig. 4 Urnochitina iirna, Eisenack 1 934. PC 1 07, chain of 3 individuals, lateral view, x 220. LV 1 0/492/9. (N34). 

 Fig. 5 Margachitina elegans Taugourdeau & de Jekhowski 1960. PC 108, lateral view, x 300. LV 1 2/494/8. (N25). 

 Fig. 6 Sphaerochitina sphaerocephala (Eisenack 1932). PC 109, lateral view, x 700, Arg5AySB7/4. 



Fig. 7 Fungochitina kosovensis Paris & Kri'z 1984. PC 1 10, LP19/SB5/57; 7a, lateral view, x 200: 7b. basal margin detail, x 1250. 

 Fig. 8 Angochitina tsegelnjiicki Paris & Grahn 1996. PC 1 1 1 . lateral view, x 200, Gerl4/468/8. (P3 1/4). 

 Fig.9 Cwgw/oc/!/7/;!i7 .serrate (Taugourdeau & Jekhowski 1960). PC 1 12. LV12/494/8. (N24), 9a, chain of II individuals, lateral view, x 130; 9b, detail of 



connection, x 500. 

 Fig. 10 Angochitina echinata Eisenack 1931. PC 113, LP8/CSB1/1; 10a, lateral view, x 350: 10b, neck detail, x 750. 

 Fig. 11 Angochitina thadeui Paris 1981. PC 1 15, LV5a/CSBl/8: 11a, lateral view, x 200; lib. apertural detail, x 650. 



Fig. 12 Angochitina ambrosi amhrosi Schweineberg 1987. PC 1 16. Ger8/461/8, (N38/3); 12a, lateral view, x 200; 12b, apertural detail, x 750. 

 Fig. 13 Angochitina chhtpaci Paris. Laufeld & Chlupac 1 98 1 . PC 1 1 7, lateral view, x 250. Arg 1 3/455/2, (J28/2). 

 Fig. 14 Angochitina echinata Eisenack 1 93 1 . PC 1 14. Ger9/456/7, (K3 1/1 ); 14a, lateral view, x 300; 14b, spiny ornament detail, x 850. 



