140 



M.K. HOWARTH 



1973 Polymorphites polymorphus (Quenstedt); Donovan & 



Forsey: 11, 12. 

 1980 Polymorphites polymorphus (Quenstedt); Schlatter: 84, pi. 



7, fig. 2 (from Wiirttemberg, Germany). 



RANGE. Found only in bed 555, Jamesoni Subzone; 2 specimens. 



Remarks. Although the lectotype of P. polymorphus, as validly 

 designated by Donovan & Forsey (1973: 12), is probably lost, 

 Schlegelmilch's (1976: 61) designation of a 'neotype' is not valid 

 because it is radically different in morphology from the lost lectotype. 

 That specimen (Schlegelmilch, 1976: pi. 28, fig. 3) is the original of 

 Quenstedt, 1885, pi. 30, fig. 9, and represents a round-whorled, striate 

 species of Polymorphites, which was described as P. lineatus (Quenstedt, 

 1845) by Schlatter (1980: 86). 



The relationship between P. trivialis (Simpson, 1843) and P. 

 polymorphus (Quenstedt, 1845) remains to be clarified: both have 

 wide ranges of morphological variation, and they may be synonyms. 

 P. trivialis is abundant in the lower half of the Brevispina Subzone, 

 but two examples off! polymorphus (PI. 7, fig. 3) that are identical 

 with the specimen figured by Dean et al. (1961: pi. 68, fig. 4) and 

 Schlatter ( 1980: pi. 7, fig. 2) were found in bed 555 in the Jamesoni 

 Subzone. They have broad whorls, widely spaced ribs and ventro- 

 lateral tubercles that are characteristic of the most strongly 

 ornamented forms of both species. 



Genus PLATYPLEUROCERAS Hyatt, 1867 



Platypleuroceras brevispina (J. de C. Sowerby, 1827) 



1827 Ammonites brevispina J. deC. Sowerby: 106, pi. 556, fig. 1. 



1843 Ammonites ripleyi Simpson: 11. 



1880/82 Aegoceras brevispina (J. de C. Sowerby); Wright: 361 



(1882), pi. 32, fig. 2, 3 (1880) (holotype, BM 43915, from 



Pabba, Inner Hebrides, Scotland). 

 1909 Uptonia ripleyi (Simpson); Buckman: pi. 2 (holotype, WM 



106, probably from beds 544.7-546.4). 

 1961 Platypleuroceras brevispina (J. de C. Sowerby); Dean et 



al.: pi. 69, fig. 1 (holotype). 



Range. From beds 544.6-550, Brevispina and Jamesoni Subzones; 

 83 specimens. 



Platypleuroceras obsoleta (Simpson, 1843) 



1843 Ammonites obsoletus Simpson: 23. 



1882 Aegoceras brevispina (J. de C. Sowerby); Wright: 361, pi. 



50, figs 13, 14(BMC3126). 

 1914 Uptonia obsoleta (Simpson); Buckman: pi. 92 (holotype, 



WM 157). 



Range. A single specimen in bed 544.7, Brevispina Subzone. 



Remarks. This is a single poorly preserved specimen with many 

 ribs and ventro-lateral tubercles which is not good enough to elucidate 

 the horizon of Simpson's larger and better preserved holotype. 



Platypleuroceras aureum (Simpson, 1855) PL 7, fig. 6 



1855 Ammonites aureus Simpson: 44 (non Ammonites aureus 



Young & Bird, 1822). 

 71855 Ammonites tenuispina Simpson: 69 (the holotype is lost - 



seeHowarth, 1962: 111). 

 1909 Platypleuroceras aureum (Simpson); Buckman: pi. 3 



(holotype, WM 107, from bed 546.2 or 546.5). 



Range. Beds 546. 1-546.5, Brevispina Subzone; 23 specimens. 



Remarks. P. aureum is a more evolute species than P. brevispina, 

 and is bituberculate (ie. both umbilical and ventro-lateral tubercles are 

 well developed). A small, typical specimen is figured in PL 7, fig. 6. 



Genus UPTONIA Buckman. 1897 



Uptonia jamesoni (J. de C. Sowerby, 1827) 



1827 Ammonites jamesoni J. de C. Sowerby: 105, pi. 555, fig. 1. 



71855 Ammonites ignotus Simpson, 1855: 61. 



71910 Uptonia ignota (Simpson); Buckman: pi. 21 (holotype, 



WM 159). 

 1973 Uptonia jamesoni (J. de C. Sowerby); Donovan & Forsey: 



12, pi. 4, fig. 3 (neotype, BM C.40426, designated by 



Donovan & Forsey, 1973: 12, from Pabba, Inner Hebrides, 



Scotland). 



Range. Beds 550-560.3, Jamesoni Subzone; 23 specimens. 



REMARKS. Many of the 23 specimens in the Jamesoni Subzone are 

 typical examples of the species, but they are mostly fragmentary, and 

 none are preserved well enough to be figured. 



Uptonia lata (Quenstedt, 1845) 



1 845 Ammonites jamesoni J. de Sowerby, var. latus Quenstedt: 

 88, pi. 4, fig. 1 (holotype, from Wiirttemberg, Germany). 



1980 Uptonia lata (Quenstedt); Schlatter: 1 13, pi. 12, figs 3, 4 

 (from Wiirttemberg, Germany). 



Range. Beds 558 and 560.3, Jamesoni Subzone; 6 specimens. 



REMARKS. A more involute species than U. jamesoni, with much 

 finer ribbing; identified according to Schlatter's (1980: 1 13) interpre- 

 tation of Quenstedt's species. 



Genus TROPIDOCERAS Hyatt, 1867 



Tropidoceras futtereri Spath, 1923 PL 7, figs 10, 11 



1923a Tropidoceras futtereri Spath: 8. 



1928 Tropidoceras futtereri Spath; Spath: 228, pi. 16, fig. 8 



(holotype, BM C.23687, from bed 118b, Charmouth, 



Dorset). 



Range. Single specimens in beds 560.3 (top) and 568 (base), 

 Masseanum Subzone. 



Tropidoceras masseanum (d'Orbigny), var. rotundum 

 (Futterer, 1893) 



1 893 Cycloceras masseanum (d'Orbigny), var. rotundum Futterer: 

 330, pi. 12, figs 3, 4 (holotype, from Wiirttemberg, Ger- 

 many). 



1980 Tropidoceras masseanum (d'Orbigny) rotundum (Futterer); 

 Schlatter: 138, pi. 19, fig. 4. pi. 20, figs 1, 2 (from 

 Wiirttemberg, Germany). 



Range. Found only at the boundary of beds 567 and 568, Masseanum 

 Subzone; 1 1 specimens. 



Genus PARINODICERAS Trueman, 1918 



Remarks . Although considered by Spath ( 1 93 8 : 8 1 ) to be a subgenus 

 of Liparoceras, Parinodiceras (including its synonym Platynoticeras) 

 is now thought to have been derived from Polymorphites and is there- 

 fore placed in the family Polymorphitidae (Donovan, 1981: 111, 138). 



