STRATIGRAPHY AND TRILOBITE FAUNAS FROM THE CAMBRIAN BURJ FORMATION 



141 



Kingaspis campbelli (King, 1923) 



Figs 21-26 



Figs 19, 20 Onaraspis palmeri (Parnes), from King's (1923) locality 83, 

 Wadi Rimeileh. 19. pygidium, external mould, Sedgwick Museum 

 A. 59463. 20, librigena, the infill between dorsal surface and doublure 

 shows white, Sedgwick Museum A. 59464. Both unwhitened. x 1.3. 



accept the generic reference but the observed differences are prob- 

 ably of specific significance: for example O. palmeri has a longer 

 frontal area and the pleural field of the pygidium tapers backwards 

 less strongly. 



King's material from Wadi Rimeileh (Figs 19, 20) consists of a 

 large librigena, some pleural fragments and two pygidia, of which 

 one is complete. The specimens are fairly well preserved in siltstone 

 and, though somewhat flattened, show granulose sculpture. The 

 pygidium appears to agree with that of O. palmeri but is longer in 

 proportion than that of O. somniurna, though it may be more similar 

 to Opik's (1968: 159) Onaraspis sp. A. 



HORIZON. Parnes (1971: pis 3, 4) recorded O. palmeri from the 

 lower and upper parts of the Timna Formation, respectively from the 

 upper part of the Hakhlil Member at Timna and from beds at Har 

 'Amram that are correlated with the Mikhrot Member. The specimen 

 from Wadi Rimeileh is from a siltstone interval in the Burj Forma- 

 tion, but its stratigraphical relationship to other localities is unknown. 



Superfamily ELLIPSOCEPHALOIDEA Matthew, 1887 

 Family ELLIPSOCEPHALIDAE Matthew, 1887 

 Subamily ELLIPSOCEPHALINAE Matthew, 1887 



Genus KINGASPIS Kobayashi, 1935 



Type SPECIES. Anomocare campbelli King, 1923, by original 

 designation. 



DISCUSSION. Geyer( 1990b: 102 (placed Kingaspis close to Ellipso- 

 cephalus, regarding the separate family (or subfamily) Kingaspididae 

 (or Kingaspidinae) as superfluous. He discussed the type species, K. 

 campbelli (King) and described several other species of Kingaspis 

 and of the closely related genus Kingaspidoides Hupe, 1953. All 

 species of these genera have features of the cranidium effaced on the 

 external surfaces, but axial and glabellar furrows are generally 

 visible on internal moulds. The weakness of these furrows makes 

 measurements taken from them rather imprecise, hampering mor- 

 phological comparisons, but the form of the glabella, which has 

 concave sides, expanded anterolateral corners and 4 or 5 pairs of 

 furrows, is consistent. 



1923 Anomocare campbelli King: 511, figs 3, 4 [described, 

 illustrated with line-drawings]. 



1935 Kingaspis campelli (sic) (King); Kobayashi: 196, pi. 23, 

 figs 9, 10 [assigned to Kingaspis; poor photographs of 

 King's specimens, the cranidium is incorrectly restored]. 



1990b Kingaspis campbelli (King); Geyer: 104, pi. 15, fig. 1 1, pi. 

 17. figs 8-10 [Moroccan specimens described and dis- 

 cussed, with full synonymy]. 



Material. King's syntype material is preserved in the Sedgwick 

 Museum, Cambridge, and from that the cranidium SM A. 1311, 

 figured by King (his fig. 3), is here selected as lectotype (Figs 21a- 

 c herein). Topotypic material is held in other museums (Hebrew 

 University of Jerusalem; Senckenberg Museum, Frankfurt; Natural 

 History Museum, London); more recently Dr J. H. Powell collected 

 further specimens, one of which is figured here (Fig. 25). 



DISCUSSION. Topotypic material is well preserved and agrees with 

 Geyer's description. He remarked that K. campbelli has a relatively 

 wider cranidium (1.3 times the length) than other species of the 

 genus. There is slight variation in transverse convexity and in the 

 evenness of the curvature in sagittal section. 



Geyer excluded the pygidium described by King from the species, 

 but because it is the only kind of pygidium found with monospecific 

 associations of the cranidia of K. campbelli. King probably origi- 

 nally associated them correctly. King's large paralectotype is shown 

 here (Fig. 22). A smaller well preserved example (Fig. 26) is 5.0 mm 

 long and estimated to have been 13 mm wide; axis more than a 

 quarter of the total width, convex, with 3 axial rings and a trace of a 

 fourth; pleural regions with 4 pleural furrows and three interpleural 

 grooves; border narrows slightly backwards. 



Originally described from the Burj Formation at Wadi Zarqa 

 Ma'in, topotypic specimens have since been figured many times 

 (see Geyer 1990b: 104). 



HORIZON. At Wadi Zarqa Ma'in K. campbelli is in earliest Middle 

 Cambrian beds associated with the lowerAmgan species Palaeolenus 

 an.ti.quus (Chernysheva), discussed below. In Morocco it is question- 

 ably recorded from the frequens Zone. 



Genus KINGASPIDOIDES Hupe, 1953 



TYPE SPECIES. Kingaspidoides armatus Hupe, 1953, by origianl 

 designation. 



DISCUSSION. Kingaspidoides was originally instituted for kingasp- 

 idids with an occipital spine (Hupe 1953), but Geyer (1990b: 1 10) 

 emended the diagnosis and distinguished the genus from Kingaspis 

 by the relatively uneven convexity of the cranidium in transverse 

 section. 



Kingaspidoides cf. obliquoculatus Geyer, 1990 Figs 27, 28 



cf. 1990b Kingaspidoides obliquoculatus Geyer: 118, pi. 18. figs 1- 

 15. 



Material. Three cranidia from Al Abrash. Ghor-es-Safi. BM 

 In24056-8. collected by Dr A. E. Day in 1909. Associated with 

 Psiloria dayi Cooper ( 1976: 283), for which this is the type locality. 



DESCRIPTION AND DISCUSSION. The glabella has independent trans- 

 verse convexity (Fig. 28e). Interocular area of fixigena scarcely 

 convex, slopes down to palpebral lobe which is short, the posterior 

 end lying well forward of SO. Glabella relatively narrow; at a line 



