Xi<C^sis''- O 



Bull. nut. Hist. Mils. Loud. [Geo\.]51(\):29-%2 X\K^ — ' ' Issued 2X Jimt- 2001 



A revision of the English Wealden Flora, III: 

 Czekanowskiales, Ginkgoales & allied 

 Coniferales 



JOAN WATSON & SUSANNAH J. LYDON 



Department of Eartli Science. Willianhson Building, The University. Manchester Ml 3 9PL 



NICOLA A. HARRISON 



Atos Origin. Wilton Centre Annexe, PO Box 54, Wilton, Middlesbrough. TS90 8JA, UK 



CONTENTS 



Introduction 30 



Geological occurrence 30 



Methods 30 



Systematic considerations 31 



Systematic descriptions 32 



Gymnospermae 32 



Order Czekanowskiales 32 



Genus Czekanowskia Heer 32 



Czekanowskia anguae sp. nov 32 



Genus Phoenicopsis Heer 34 



Phoenicopsis rincewindii sp. nov 36 



Order Ginkgoales 39 



Genus Ginkgoites Seward 39 



Ginkgoites weatherwaxiae sp. nov 40 



Ginkgoites nannyoggiae sp. nov 45 



Ginkgoites garlickianus sp. nov 47 



Ovule attributed to Ginkgoites weatherwaxiae 5 1 



Order Coniferales 55 



Incertae sedis (family uncertain) 55 



Genus Pseudotorellia Florin 55 



Pseiidotorellia linkii (Romer) 56 



Pseudotorellia vimesiana sp. nov 61 



Genus Sciadopityoides Sveshnikova 63 



Sciadopiryoides greeboana sp. nov 63 



Genus Sulcatocladiis Watson & Harrison 69 



Sulcatocladits robiistiis Watson & Harrison "70 



Sulcatocladiis dihbleri sp. nov ^ ^' 



? Family Taxaceae ; '^ 



Genus Torreyites Seward '-^ 



Torreyites detriti sp. nov j '3 



Palaeoecology '" 



Acknowledgements ' 



References ° 



Appendix: List of specimens studied "^ 



Synopsis. Eleven gymnosperm leaf species are identified and described from the English Wealden llora, attributed to the orders 

 Czekanowskiales. Ginkgoales and needle-leaved Coniferales. All occur as fragments in the "plant debris beds" which are common 

 throughout the Wealden succession in East Sussex. Dorset and the Isle of Wight. Two new species assigned to the Czekanowskiales 

 are placed in the genera Czekanowskia Heer and Phoenicop.m Heer; three new species assigned to the Ginkgoales are placed in 

 the form-genus Ginkgoites Seward; an ovule containing monosulcate pollen grains, associated with one of the Ginkgoites leaves 

 is described and illustrated. Needle-leaved conifers of uncertain family attribution comprise two species assigned to the genus 

 Pseudotorellia Florin, one species to Sciadopityoides Sveshnikova. The shoot species Sulcatocladiis robustus Watson & Harrison 

 is known to be attributable to the leaf species Pseudotorellia linkii (Romer); a second species of Sulcatocladiis Watson & Harrison 

 is attributed to the new species of Sciadopityoides. A leaf species with stomata in two grooves is referred to the form-genus 

 Torreyites Seward and tentatively to the family Taxaceae. The Czekanowskiales are recorded from the Lower Cretaceous of 

 western Europe for the first time and the palaeoecological implications of these additions to the English Wealden flora are 

 discussed. 



©The Natural History Museum. 2001 



