48 



J. WATSON, S.J. LYDON & N.A. HARRISON 







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Fig. 14 A-D Ginkgoites nannyoggiae sp. nov. A-C show the holotype from Fairlight CLis s tai.ies of Ashdown Beds, Galley Hill, Sussex. D from Wessex 

 Formation, Mupe Bay, Dorset. A, upper cuticle showing oval scar overlapping 3 ordmary epidermal cells V 64545, LM, x 500: B, inside of upper cuticle 

 showing pitting, V.64545, SEM, x 1000; C, oval scar on outside of upper cuticle, V.64545, SEM, x 750; D, possible post-mortem bite-mark, V.64546, 

 LM, X 100. 



garlickianus sp. nov. figured here have been found as dispersed 

 fragments with good cuticle preservation in the 'Grange Chine Black 

 Band' plant debris bed at Grange Chine on the South West coast of 

 the Isle of Wight (Locality Lll of Stewart 1978). This bed, as 

 previously discussed, is an important source of vertebrate material 

 (see introduction). Oldham (1976) described and figured material 

 belonging to this species from various Wessex Formation localities 

 along the South West coast of the Isle of Wight. Stratigraphical 

 range: Barremian. 



Description and discussion. Ginkgoites garlickianus sp. nov. is 

 one of three species from the English Wealden newly described in 

 the present work and attributed to the genus Ginkgoites Seward. 

 Although Oldham (1976) recognized the cuticle of G. garlickianus 

 as of ginkgoalean affinity, he did not place it within a genus or erect 

 a species for it, using instead the Biorecords code 26 Gink GiA. 



The few leaf fragments of G. garlickianus recognized so far 

 indicate a lamina which is deeply divided into narrow segments and 

 Fig. 16A shows a leaf segment with its complete width intact. 



However, no conclusive evidence from a petiole or branching or leaf 

 apices of segments, which would establish a gross morphology for 

 the leaf of this species, has yet been recognised. 



The cuticle of the upper epidermis (Figs 16C-G; 17B; 18A-C) is 

 covered by numerous, prominent papillae and trichomes on its outer 

 surface (Figs 16B, C) and is thus easily distinguished from that of the 

 lower epidermis (Figs 16B; 19A-F) which has a smooth and feature- 

 less outer surface (Fig. 19B). The upper cuticle shows the ordinary 

 epidermal cells arranged in longitudinal files (Fig. 16C, E); this 

 combines with the papillate nature of the cells to provide a ridged 

 structure to many areas of cuticle (Fig. 16D, F). The trichomes, 

 which are up to 50 pm long, are often flattened (Fig. 16E) and only 

 broken bases remain in some cases. The stomata are randomly 

 arranged and usually longitudinally orientated (Figs 16E; 17B, 18A- 

 C), and are similar to those of the lower surface. Although easily 

 recognised in the SEM, stomata are less easy to identify in the light 

 microscope, especially at low magnifications. 



The ordinary epidermal cells of the lower cuticle are mostly 

 isodiametric and randomly arranged (Figs 19A, C); the outer surface 



