50 



J. WATSON, S.J. LYDON & N.A. HARRISON 







H'i. 









\mmm 



WM% 



'Wi 



"*'"^ 







WSBmfT^ 



tWHElJ 



1 kVJ^^^^^^^^^^^^^I 





HUKfa A V 



h| 







p 



1 



iwm 





1 



ii 





Eu^ 



iH 



iM^fl 



B^^^^P^ 



iS , ^ ' ilJK^; .•' 



Fig. 16 A-G Ginkgoites garlickianus sp. nov. A, holotype; leaf fragment showing full segment width. V.64548, LM, x 10: B, segment margin of holotype, 

 dividing upper (left) and lower (right) cuticle, V.64548. LM, x 75; C-G upper cuticle; C, heavily papillate cuticle, showing large ridge on left. V.64548, 

 LM, X 125; D, outer surface of cuticle showing whole and broken papillae. V. 64549, SEM, x 125; E, inside of cuticle showing ordinary epidermal cells in 

 rows and stomatal distribution, V.64548. SEM, x 125; F, ridged outer surface bearing flattened papillae and possible stomatal opening (bottom right), 

 V.64549, SEM, x 500; G, single flattened papilla, V.64549, SEM, x 1000. 



knowledge of this species. This is a distinct possibility since its 

 distinctive suite of characters makes for fairly easy recognition, but 

 it requires further intensive searching of debris material. 



Comparison. Of the 3 species of Ginkgoites present in the English 

 Wealden, the most easily distinguished is Ginkgoites nannyoggiae 

 which has much the widest leaf segments, stomata confined to the 

 densely papillate lower surface, and an upper cuticle which shows 

 isodiametric cells only. Both Ginkgoites garlickianus and Ginkgoites 

 weatherwaxiae have narrow segments, stomata on both surfaces and 



an upper cuticle with more elongate ordinary epidermal cells than 

 the lower. However, G. garlickianus has a heavily papillate, ridged 

 upper cuticle, whereas that of G. weatherwaxiae is generally smooth. 

 The stomata of the lower surface of G. weatherA'axiae are oval in 

 outline and typically have 6 papillate subsidiary cells. Those of G. 

 garlickianus are more circular in outline with a smaller opening and 

 typically have 5 subsidiary cells on which the presence of papillae 

 varies considerably between specimens. The cuticle of G. 

 weatherwaxiae is in some ways more similar to that of Czekanowskia 

 anguae which is also present in the plant debris beds of Worbarrow 



