58 



J. WATSON, S.J. LYDON & N.A. HARRISON 



1852 Pinites Linkii (Romer) Ettingshausen: 27. [Name change 



only] 

 1 87 1 Abietites Linkii pro parte (Romer), emend. Schenk: 24 1 , pi. 



39; pi. 40, figs 1-5, 7. [Northwest Germany] 

 1 923 Podozamites sp. of. affinis (Schenk) Lipps: 356, text-fig. 25. 



[Northwest Germany] 

 1926 Pityophyllum crassum Seward: 106, text-fig. 16. [Western 



Greenland] 

 1936 Abietites Linkii (Romer); Michael: 61, pi. 3, figs 1, 2. 



[Northwest Germany] 



1960 ror(?///fl-aehnliche Daber: 606, pi. 15, figs 3a, 5. [Wilsnack 

 borehole. Northeast Germany] 



1961 Abietites linkii (Romer); Benda: 624, pis 39-41. [North- 

 west Germany] 



1969 Pseudotorellia heterophylla Watson: 248, pi. 6. figs 6, 7; 

 text-figs 59-64. [England] 



1969 Tritaenia linkii (Romer) Magdefrau & Rudolf: 296, text- 

 figs 1-5. [Northwest Germany] 



1976 24 GINK ToB Oldham [Code used instead of Linnean 

 binomial]: 458, pi. 68, figs 1-3. [England] 



1976 25 GINK ToA Oldham [Code used instead of Linnean 

 binomial]: 458, pi. 68, figs 4-8. [England] 



1980 Pseiidotorellia-a.&hn\\c\\c Daber: 276, pi. 106, fig. 3; text- 

 fig. 109. [Gorlosen and Wilsnack boreholes. Northeast 

 Germany] 



1984 Abietites //wA// (Romer); Van der Burgh & Van Konijnenburg- 

 Van Cittert: 390, pi. 9, fig. 1. [Northwest Germany] 



1991 Tritaenia linkii (Romer) sensu Magdefrau & Rudolf; Bose 

 & Manum: 14. 



1991 Tritaenia crassa (Seward) Bose & Manum: 15, fig. 6. 

 [Western Greenland] 



1991 Tritaenia linkii (R5mer); Wilde: 366, figs 1-3. 



1996 Abietites linkii (Romer); Watson & Alvin: 9. [Name in list]. 



1996 Pseudotorellia heterophylla Watson; Watson & Alvin: 9. 

 [Name in list]. 



1 998 Pseudotorellia linkii ( Romer) Watson & Harrison: 24 1 , figs 

 lA, B; 3; 4A-J; 5-10; 1 1 A-G; 12-15; 16D; 20A, B. 



2000 Tritaenia linkii (Romer); Manum, Van Konijnenburg-Van 

 Cittert & Wilde: 262, fig. 2; pi. 1 , figs 1,2.6; pi. 2, figs 1 , 3, 

 4. 



Tritaenia crassa (Seward); Manum, Van Konijnenburg-Van 

 Cittert & Wilde: 263, pi. 1, fig. 5; pi. 2, fig. 2. 



Diagnosis. See diagnosis of Pseudotorellia linkii (Romer) given 

 by Watson & Harrison (1998: 242) which requires no emendation at 

 present. 



Neotype and type locality. Specimen 1997/102, Museum fiir 

 Naturkunde, Berlin. Collected Gothan 1940, Osterwald, northwest 

 Germany; selected and figured by Watson & Harrison ( 1 998, figs 5C, 

 7E, F). 



Material and occurrence. Pseudotorellia linkii (Romer) ( 1 839) 

 in the needle-leaved form occurs in great abundance at some localit- 

 ies in the Wealden of northwest Germany (Wilde 1991; Watson & 

 Harrison 1998; Manum £>?«/. 2000) with ellipticalleaves in Germany 

 occurring as a rare hand specimen and in boreholes. In the English 

 Wealden P. linkii occurs mainly from debris deposits but also as 

 single needles on some of the hand specimens collected by Rufford. 

 In the Weald Basin it has been isolated from plant debris beds in the 

 Ashdown Beds Formation at Ecclesbourne Glen and Fairlight where 

 needles and elliptical leaves have been found in the same samples 

 (Watson 1964, 1969), and Oldham (1979) recorded the species from 

 Haddock's Rough and Galley Hill. In the Wessex Basin much more 



Fig. 23 A-G Pseudotorellia linkii (Romer) Watson & Harrison. A, B, 

 wide elliptical leaves from the English Wealden. A, V.51525; B, 

 V.5 1 528; C, leaf showing two dichotomising veins and positions of 

 stomata, V.64243. x 15; D, part of sub-parallel-sided leaf with 4 veins, 

 V.51527; A, B, D after Watson 1969, all x 10; E, distal part of leaf with 

 3 resin canals, V.64239. x 10; F. distal leaf fragment with 3 resin canals, 

 V.64240, X 10: G, leaf with twisted, wrinkled base; i, lower surface 

 showing position of 3 stomatal bands: ii, upper surface with grooved 

 midline. V.64250, x 6. 



extensive material has recently been recovered from several plant 

 debris horizons in the 'lignite beds' of Arkell (1947) at Worbarrow 

 Bay, Dorset. P. linkii has also been recognised in the Lower Creta- 

 ceous flora of Angiarsuit, western Greenland (Seward. 1926). The 

 specimens shown in Figs 22 and 23, from England, Germany and 

 Greenland, are all housed in the NHM (see Appendix). 



Description. The extensive study of Pseudotorellia linkii (Romer) 

 by Watson & Harrison (1998) involved bulk maceration of many 

 samples from both England and Germany with the isolation of large 

 numbers of individual leaves displaying a considerable range and 



