REVISION OF THE ENGLISH WEALDEN FLORA 



65 



H 



Fig. 27 A-O Sciadopityoides greeboana sp. nov. All dispersed portions of leaves from the Fairlight Clay at Galley Hill. East Sussex. A-G, I-N, apical 

 fragments with lower side uppermost to show position of stomatal band (dashed) and resin canals (dotted); Hi, ii, lower and upper surfaces of proximal 

 portion of leaf with constricted base; A-N, V.64572-64585. all x 15; O, distribution of stomata in a band on the lower cuticle. ^64586. x 50. 



also common, extending the length of one or more cells in a fi le ( Figs 

 28B; 29B), less commonly, hollow dome-like papillae are present 

 (Fig. 29C). Beneath the thickly cutinized outer periclinal walls of the 

 epidermal cells, a thin layer of cuticle representing the inner periclinal 

 wall is frequently developed (Fig. 29D). The outlines of the hypoder- 

 mal cells are also usually preserved as thin sinuous strands of cuticle 

 (Figs 28B; 29E, F). The hypodermal cells are commonly twice the 

 width of the epidermal cells and often have their longitudinal walls 



directly beneath those of the epidermal cells. They are. however, 

 shorter than the overlying epidermal cells and the end walls of the 

 two cell layers are rarely in line. 



On the lower leaf surface, the lateral, stomatal-free /ones are 

 identical to the upper cuticle. The stomatal band with its thinner 

 cuticle is quite distinct and has clearly defined edges. Hypodermal 

 cutinization is completely lacking. The ordinary epidermal cells in 

 the band are also rectangular and arranged in tiles (Fig. 29A-D). 



