Bull. nat. Hist. Miis. Land. (Geol.) 55(2): 81-137 Issued 25 November 1999 



Late Cretaceous-early Tertiary echinoids from 

 northern Spain: implications for the 

 Cretaceous-Tertiary extinction event 



A. B. SMITH 



Department of Palaeontology. The Natural History Museum. Cromwell Road. London SW? 5BD 



J. GALLEMI 



Museo de Geologia. Pare de la Ciutadella s/n. 08003 Barcelona. Spain 



C. H. JEFFERY 



Departments of Palaeontology and Zoology. The Natural History Museum. Cromwell Road. London SW7 5BD 



G. ERNST 



Institut fiir Paldontologie der Freien Universitat. Malteserstrafie 74-100. Haus D, D 12249 Berlin 33. Germany 



P. D. WARD 



Department of Geological Sciences, AJ-20, University of Washington, Seattle 98195, USA. 



CONTENTS 



Synopsis 82 



Introduction 82 



Geological Setting 83 



Details of Sections 83 



The Santander Region 83 



The Basque Region 84 



The Navarra Region 86 



Faunal Changes across the Cretaceous-Tertiary Boundary 87 



Systematic Descriptions (Smith, Gallemi & Jeffery) 89 



Order CidaroidaClaus, 1880 89 



Family Psychocidaridae Ikeda, 1936 89 



Genus 7>/oarfamPomel, 1883 89 



Family Cidaridae Gray, 1825 91 



Genus Temnocidaris Cotteau. 1863 91 



Indet. cidarid plates 92 



Cohort Echinacea Claus, 1876 92 



Family Orthopsidae Duncan, 1889 92 



Genus Orthopsis Coneau, 1864 92 



Order Calycma Gregory, 1900 92 



Family Saleniidae Agassiz, 1838 92 



Genus Salenia Gray, 1835 92 



Order Arbacioida Gregory, 1900 93 



Family Goniopygidae Smith & Wright, 1993 93 



Genus Goniopygiis L. Agassiz, 1838 93 



Order Phymosomatoida Mortensen, 1904 94 



Family Phymosomatidae Pomel. 1883 94 



Genus GoMrtfena Lambert, 1888 94 



Genus O/p/otogwa Schluter, 1870 96 



Genus Phymosoma Haime, in d'Archiac & Haime, 1853 96 



Genus Aca/i//!ec/!/>7M^ Duncan & Sladen, 1882 96 



Cohort Irregularia Latreille, 1825 97 



Order Holectypoida Duncan, 1889 97 



Family Holectypidae Lambert, 1 899 97 



Genus Coenholectypus ?ome\, 1883 97 



Genus Camerogaterus Qusn^XtAU 1873 97 



Order Echinoneoida Clark, 1925 97 



Family Conulidae Lambert, 191 1 97 



Genus Comdus Leske, 1778 97 



Gtn\is Adelopneusles GmXhKT. 1889 100 



Order Cassiduloida Claus, 1880 '00 



© The Natural History Museum, 1999 



