82 A.B. SMITH ETAL. 



Family Pygaulidae Lambert, 1905 100 



Genus Echino^olerus Konig, 1825 100 



Family Echinolampadidae Gray, 1851 101 



Genus Gitolampas Gauthier, 1889 101 



Family Clypeolampadidae Kier, 1962 102 



Genus Clypeolampas Pomel, 1869 102 



Unnamed Family 102 



Genus Nudeopygus L. Agassiz. 1840 102 



Family Cassidulidae Agassiz & Desor, 1847 104 



Genus Oolopygiis d'Orbigny, 1856 104 



Genus /?/!y«f/!o/;vgi«d"Orbigny, 1856 104 



Genus Rhyncholainpas Agassiz, 1869 105 



Family Faujasiidae Lambert, 1905 105 



Genus Zujfardia Checchia-Rispoli, 1917 105 



Order Holasteroida Durham & Melville, 1957 106 



Family Holasteridae Pictet, 1857 106 



Genus Ojfaster L.Kgassxz. 1836 106 



Genus Cardiaster Forbes, 1850 106 



Genus Hemipneustes Agassiz, 1836 107 



Family Echinocoridae Lambert, 1917 108 



Genus Echinocorys heske, 1778 108 



Genus Jeronia Seunes, 1888 109 



Family Stegasteridae Lambert, 1917 1 10 



Genus Stegaster Pomel 1883 110 



Genus Tholasier Seunes. 1891 114 



Genus Pseudojf aster Lambert, in Lambert & Thiery, 1924 1 18 



Genus Gatemer Seunes, 1889 118 



Order Spatangoida Glaus, 1876 118 



Family Micrasteridae Lambert 1920 118 



Genus Micraster Agassiz, 1836 119 



Genus Cyclaster Cotteau, in Leymerie & Cotteau, 1856 120 



Genus Isaster Desor, 1858 124 



Family Hemiasteridae Clark, 1917 124 



Genus Hemiaster Agassiz, in Agassiz & Desor, 1847 125 



Family Corasteridae Lambert, in Lambert & Thiery, 1924 126 



Genus Coroifer Cotteau, 1886 126 



Genus HomoeasterPomeX, 1883 130 



Genus Ovulaster Coneau, 1884 130 



Family Schizasteridae Lambert, in Doncieux, 1905 130 



Genus Linthia Desor, 1853 130 



Family Aeropsidae Lambert, 1896 131 



Genus Sphenaster Jeffery 131 



References 132 



Appendix 135 



Synopsis. The Maastrichtian and Palaeocene echinoid faunas are described from Santander, Province of Cantabria, the Basque 

 region of Spain and France, and the region around Irurtzun, Province of Navarra. These deposits range from shallow shelf 

 carbonates to upper continental slope elastics. A total of 58 species are described, 40 from the Maastrichtian. nine from the Danian 

 and nine from the Thanetian. Echinoid extinction is estimated to be around 40% in shallow water settings, but may have been more 

 intense in deeper settings. Extinction levels are comparable with the other well-studied western European faunas of the Danish 

 basin and the Limburg region. A striking shift in the geographic ranges of taxa took place between the end of the Cretaceous and 

 the Palaeocene. Taxa migrated both from the Danish basin southwards and from the Tethyan region northwards at this time. 



The first fossil representative of the Recent deep-sea family Aeropsidae is described from the early Thanetian of Navarra. New 

 taxa described here include the genus Sphenaster and the species Tylocidaris trempinus. Camewgalerus cantabrius, Adelopneustes 

 ernsti, Stegaster palaeocenicus, Ovulaster reticidatus and Sphenaster larwnbensis. 



INTRODUCTION 



directed towards documenting the synchroneity and duration of 

 extinctions, through microstratigraphical sampling in 'complete' 



boundary sections (e.g. Marshall & Ward, 1996, MacLeod, 1998). 

 The end of the Cretaceous saw the extinction of many marine Much less emphasis has been given to the ecological and geographi- 

 invertebrates, but precisely what events brought about this extinc- cal pattern of extinction at this time, with the notable exception of 

 tion, and over how short a time interval these events took place Jablonski & Raup (1994) and Smith & Jeffery (in press). Under- 

 remain the subject of heated debate. Most research so far has been standing what, if any, environmental pattern there is to survivorship 



I 



