86 



A.B. SMITH ETAL. 



Zumaia 



i->-y) 







to 



Fig. 4 Composite stratigraphic section of tlie Maastrichtian of the Basque 

 region from Zumaia. Gipuzkoa, Spain to Bidart. Pyrenees-Atlantique, 

 France (from Ward & Kennedy 1993) showing the distribution of 

 echinoids. 



pioneering work of Seunes (1888a, 1888b and 1889). He reported 

 the occurrence of Cidaris beaugeyi Seunes, Jeronia pyrenaica Seunes, 

 Coraster beneharnicus Seunes, Echinocorys douvillei Seunes, 

 Stegaster heberti Seunes, S. bouillei Cotteau, Gibbaster munieri 

 Seunes [=Tholaster munieri (Seunes)] and Galeaster bertrandi 

 Seunes from the vicinity of Bidart. 



The Navarra region 



This has the most complicated geology of the three regions, being 

 strongly affected by strike-slip faulting. Maastrichtian deposits are 

 represented by two facies: shelf platform carbonates (studied at 

 Olazagutia Pass) and shelf basinal mudrocks (studied at Larumbe 

 andSarasate)(Fig. lb). Table I lists the echinoids collected from this 

 region. The overlying Danian and early Thanetian consist of a 

 sequence of coccolith limestones that in places includes reworked 

 pebble beds. 



Table 1 Echinoids from the Navarra district (see Fig. lb for map of area). 

 LOCALITY 1. Maastrichtian, carbonate platform facies; Olazagutia Pass. 



Tylocidaris (Sardocidaris) ramondi (Leymerie) 

 cidaroid plates indet. 



Salenia (Pleurosalenia) maastrichtensis Schluter 

 Conulus gigas (Cotteau) 

 Echinogalerus muelleri (Schluter) 

 Nucleopygus scrobicidatus (Goldfuss) 

 Offaster leymeriei Cotteau 

 Hemipneustes pyrenaicus Hebert 

 Hemipneustes striatoradiatus (Leske) 

 Hemiaster koninckanus d"Orbigny 

 Hemiaster prunella (Lamarck) 

 Cyclaster sp. 



LOCALITY 2. Late Thanetian, Casas de Oraien. 



Gauthieria pseudomagnifica (Cotteau) 



Adelopneustes ernsti sp. nov. 



Cardiaster sp. 



haster aquitanicus (de Grateloup) 



Echinocorys scutata forma cotteaui Lambert 



Micrasler tercensis Cotteau 



Hemiaster stella (Morton) 



Linthia sp. Cotteau 



LOCALITY 3. Danian-early Thanetian at Larumbe. 



Coraster vilanovae Cotteau 

 Coraster beneharnicus Seunes 

 Sphenaster larumba sp. nov. 

 Jeronia pyrenaica Seunes 

 Echinocorys sp. 



LOCALITY 4. Maastrichtian; basinal facies. 



Motorway cutting at Sarasate, just South-East of Irurzun. 



Stegaster bouillei (Cotteau) 



Stegaster altus Seunes 



Echinocorys scutata forma ovata Leske 



Ovulaster reticulatus sp. nov. 



Pseudoffaster caucasicus (Dru) 



Cyclaster sp. 



Danian near summit of hill at Sarasate 



Cyclaster aturicus (Seunes) 



Cyclaster gindrei (Seunes) 



Coraster beneharnicus Seunes 



LOCALITY 5. Danian of Erice. 



Coraster vilanovae Cotteau 



Coraster beneharnicus Seunes 



Cidaroid spines 



Echinocorys scuata forma pyrenaica Seunes 



Cyclaster grindrei (Seunes) 



LOCALITY 6. Late Danian at Aristregui. 



Stegaster paleocenicus sp. nov. 

 Jeronia pyrenaica Seunes 

 Coraster vilanovae Cotteau 

 Echinocorys scutata Leske 



Olazagutia Pass. The carbonate platform succession was studied 

 at Olazagutia Pass (Fig. lb), the type locality of the Puerto de 

 Olazagutia Formation (Amiot, 1982), along the road leading from 

 Olazti to the Urbasa Range. The units that lie at the top of the 

 Olazagutia Formation yield the richest and most varied fauna. These 

 beds were deposited in an external platform setting (Amiot, 1982). 

 Here the Maastrichtian succession commences with a massive 

 bedded limestone (Fig. 5). This is succeeded by a series of thinner 

 sandy limestones, calcareous sandstones and marls which are richly 

 fossiliferous. A thick sandy marl with a rich bryozoan fauna at the 



