LATE CRETACEOUS-EARLY TERTIARY ECHINOIDS 



randomly arranged, but occassionally form longitudinal lines near 

 the distal tip. 



Material studied. BMNH EE4545a-h, EE6265-68, MGB 



37444-49, 37480-87. 



Occurrence. The species is based on tests and isolated spines 

 from the Maastrichtian of Monleon and Gensac, Haute-Garonne, 

 French Pyrenees. In Spain spines and isolated interambulacral plates 

 are moderately common in the Maastrichtian, horizon 6, of Olazagutia 

 Pass, Navarra province. Lambert (1907: 712) recorded this species 

 from 'Garumnian facies' of Bouzin (Haute-Garonne, France) which 

 is Danian in age. This, however, is based on a test and the character- 

 istic spines of T. ramondi were not apparently collected. It is not 

 therefore certain to what species this test should be assigned. 



Remarks. A readily distinguishable species of Tylocidaris on 

 account of its characteristically shaped spines and slightly greater 

 number of interambulacral plates. The spines are readily distinguish- 

 able from those of Tylocidaris trempinus in both shape and ornament. 

 Whereas T. ramondi spines are fat, glandiform and rapidly taper to a 

 blunt point distally, those of T. trempinus are more fusiform and 

 distally elongate, tapering to a more distinct point. More impor- 

 tantly, the ornament on the shaft of T. ramondi consists of rather fine 

 and dense granulation that is arranged irregularly, whereas in T. 

 trempinus the ornament comprises rather stout and well-formed 

 beaded ribs, from the neck to the distal tip. 



Tylocidaris (Sardocidaris) trempinus Gallemi & Smith, 



sp. nov. PI. 1, fig. 2 



?1851 Spines of an unknown Cidaris,Leymerie: 192,pl.9,fig. 13. 

 ?1862a Cidaris ramondi Leymerie; Cotteau: 315 (pars), pi. 1076, 



figs 9, 13. 

 1920 Tylocidaris ramondi Leymerie; Lambert: 4. 

 1992 Sardocidaris ramondi (Leymerie); Gallemi: 56, photo 4. 



Diagnosis. Spines up to 28 mm in length, fusiform, with widest 

 part of shaft about one-third above the base. Shaft tapering to a small 

 blunt point distally and also tapering towards the base; width less 

 than one-third of length. No distinct handle is developed beneath the 

 shaft. The entire shaft is ornamented with relatively stout beaded 

 ribs, densely spaced. Base imperforate and non-crenulate. 



Types. Holotype Universitat Autonoma de Barcelona, Palaeonto- 

 logical Collections PUAB-4321 illustrated by Gallemi (1977: 11, pi. 

 1, fig. 3); paratypes PUAB-4055 {op. cit. pi. 1, fig. 4) and PUAB- 

 43626-32. 



Other material studied. 

 37514,37545. 



91 



BMNH EE61 14-15, MGB 37513. 



Occurrence. In Spain this species occurs in the Upper Campanian 

 of Toralla, near La Pobla de Segur. Tremp Basin and Lower 

 Maastrichtian. Serra dels Homes Morts (Salas de Pallars) and Orcau, 

 Tremp Basin (Gallemi, 1992). It also occurs in the Maastrichtian, 

 horizons 4 and 6, at Santander. 



Remarks. Differs from spines of T. ramondi in having an orna- 

 ment of continuous beaded ribs instead of dense irregular granulation. 

 Differs from spines ofTylocidaris clavigera (Mantell ), T. inexspectata 

 Jagt & Van der Ham and T. hemmoorensis Salah & Schmid in its 

 strongly fusiform and distally tapering shape. Spines of I haltica are 

 more bulbous and have only fine irregular granulation. Leymerie 

 ( 1 85 1) reported finding very occasional spines in the Maastrichtian 

 of Monleon and Gensac, Haute-Garonne, French Pyrenees that were 

 clearly different from those of T. ramondi. Although he did not name 

 these, they appear to match those from Tremp. 



Tylocidaris sp. PI. 1, fig. 5 



Diagnosis. Small test approximately 17 mm in diameter and 9.3 

 mm in height. Peristome and apical disc similar in size, approxi- 

 mately 5.5-6 mm in diameter. Interambulacral zones wide with 

 primary tubercles offset towards adradial margins. Primary tuber- 

 cles imperforate, noncrenulate. Six plates in each column. Ambulacra 

 weakly sinuous with primary tubercles to each plate and perradial 

 granulation. 



Material STUDIED. BMNHEE6109. 



Occurrence. Maastrichtian, horizon 2, of Santander, Cantabria. 



Remarks. The various species of Tylocidaris all have very similar 

 test morphology. Without associated spines it is virtually impossible 

 to determine such material to species level. 



Family CIDARIDAE Gray, 1825 



Genus TEMNOCIDARIS Cotteau, 1863 



Diagnosis. Cidaroids with perforate, non-crenulate tubercles; 

 some fully formed adapical plates with rudimentary tubercles. 

 Ambulacral pores non-conjugate. 



Temnocidaris (Temnocidaris) magnifica Cotteau, 1 863 



Pi. 1, figs 3, 4 



1863a Temnocidaris magnifica Cotteau. 357. p\. 1085. 1086. 



PLATE 1 



Fig. 1 Tylocidaris (Sardocidaris) ramondi (Leymerie. 1851). BMNH EE4545, Maastrichtian of Olazagutia Pass, Navarra. Spine. x2. 



Fig. 2 Tylocidaris (Sardocidaris) trempinus sp. nov.. BMNH EE61 14. Maastrichtian of Santander, Cantabria. Spine. x2. 



Figs 3, 4 Temnocidaris (Temnocidaris) magnifica Cotteau, 1 863. Maastrichtian of Santander. Cantabria. 3, BMNH EE4415. interambulacral plates and 



associated ambulacrum, x2: 4. BMNH EE6I 1 1. spine. x2. 

 Fig. 5 Tylocidaris sp. BMNH EE6109, Maastrichtian of Santander, Cantabria. Test in profile, x2. 



Fig. 6 Orthopsis miliaris (d' Archi&c, 1835), BMNH EE61 18. Maastrichtian of Santander. Cantabria. Apical view of test. xl.5. 

 Fig. 7 Indet. cidarid. BMNH EE61 13. Maastrichtian of Santander, Cantabria. Interambulacral plate and associated ambulacral plates. x2. 

 Figs 8-10 Salenia (Pleurosalenia) maestrichtensis Schliiter. 1892. BMNH EE4423. Maastrichtian of Olazagutia Pass. Navarra. Apical, lateral and oral 



views. x2. 

 Figs 11, 12 Acanthechinus sp. BMNH EE6128. Maastrichtian of Santander. Cantabria. Lateral and apical views. x2. 



Figs 13, 14 Salenia (Pleurosalenia) scabra (Nestler. 1965). BMNH EE6I 10, Maastrichtian of Santander, Cantabria. lateral and apical views, xl.5. 

 Fig. 15 Phymosoma hexaporum Lambert, 1927, MGB 37556, Maastrichtian of Santander, Cantabria. Adoral portion of test, x2. 

 Figs 16, 17 Goniopygus tetraphyma Lambert, 1907, BMNH EE44I9. Maastrichtian of Santander. Cantabria. Lateral and apical views, x2. 

 Figs 18-20 Gauthieria pseudomagnifica (Cotteau, 1877), BMNH EE61 17, Thanetian of Casas de Oraien, Navarra. Apical, lateral and oral views. 



