92 



A.B. SMITH ETAL. 



non\9%2 Temnocidaris magnifica Salah: 209, pi. 1, figs 1-3 

 [=Stereocidaris serrate Desor] 



Diagnosis. Large Temnocidaris with relatively fine extra- 

 scrobicular tuberculation, six abreast adradially and up to 16 abreast 

 interradially; interambulacra with ca. seven plates in a column; 

 plates almost twice as wide as tall, so that primary tubercles occupy 

 virtually the entire plate height and scrobicular circles are tangential 

 at the ambitus. Ambulacral plates with large primary tubercle, fol- 

 lowed by one or two secondary tubercles and then more irregular 

 perradial tuberculation in two rows. Pitting strongly developed both 

 in ambulacra and in interambulacral extrascrobicular zones. Spines 

 long, cylindrical with finely thomed ribs. 



Occurrence. Maastrichtian, horizons 2 and 7/8, of Santander. 

 The species was first described from the Maastrichtian of Aurignac, 

 Haute-Garonne, France, and is also known from Cotentin, south 

 west France. 



Material studied. BMNH EE4415, EE61 11. 



Remarks. This species is distinguished from the very closely 

 related T. danica by its finer extrascrobicular granulation. In T. 

 magnifica extrascrobicular granulation is about 67% the diameter of 

 that in T. danica. Only isolated spines were found in beds at Santander 

 which are tentafively assigned to this species because of their simi- 

 larity to spines of other species of T. (Temnocidaris). 



Indet. cidarid plates 



Occurrence in spain. Maastrichtian, horizons 2, 4, 5, and 7/8 of 

 Santander; Maastrichtian. horizon 6, of Olazagutia Pass. 



Remarks. Isolated cidaroid plates with perforate noncrenulate 

 tubercles and narrow extrascrobicular tuberculation and belonging 

 to at least two different taxa have been collected from the 

 Maastrichtian of Santander and Olazagutia. This material is too 

 fragmentary to place even to genus. 



Cohort ECHINACEA Claus, 1876 



Diagnosis. Regular echinoids with keeled teeth and compound 

 ambulacral plating. 



Family ORTHOPSIDAE Duncan, 1889 

 Genus ORTHOPSIS Cotteau, 1864 



Diagnosis. Primary tubercles perforate and non-crenulate. 

 Ambulacra trigeminate with acrosaleniid-style plate compounding. 



Orthopsis miliaris (d'Archiac, 1835) 



PI. I, fig. 6 



1835 Cidarites miliaris d'Archiac: 170, pi. 11, fig. 8. 



1 864a Orthopsis miliaris (d' Archiac); Cotteau, in Cotteau 1 86 1 a- 



67a:558,pl. 1131, figs 1-16 (with full synonymies up to the 



date). 

 1 895 Orthopsis morgani Cotteau & Gauthier: 87, pi. 1 4, figs 6-9. 

 1985 Orthopsis miliaris (d'Archiac); Geys; 134, pi. 5, figs 8-10. 

 1991 Orthopsis miliaris (d'Archiac); Smith & Bengtson; 30, 



text-fig. 23, pi. 8, figs B-F. 

 1995 Orthopsis miliaris (d'Archiac); Smith: 136, figs 12-14, pi. 



2, figs 4-5; pi. 3, figs 1-9. 



Diagnosis. Test up to ca. 25 mm in diameter; depressed; rounded 

 in profile with ambitus slightly below mid-height. Apical disc dicycUc; 



usually preserved in position. Ambulacra straight with uniserial 

 columns of pore-pairs. Plating trigeminate with all three elements 

 reaching the perradial suture; primary tubercle overlapping two of 

 the three elements only. Interambulacra with central primary tuber- 

 cle and slightly smaller secondary tubercles on adradial and interradial 

 margins from ambital region adorally. Peristome moderately large 

 and slightly sunken. 



Occurrence in spain. Maastrichtian, horizons 7/8 and 9, of 

 Santander; Upper Campanian of Barranc de Vilanova, Toralla, near 

 La Pobla de Segur, Tremp Basin and Lower Maastrichdan of Homes 

 Morts Member, Salas de Pallars, Tremp Basin (Gallemi, 1992). This 

 species is widely distributed in the Upper Cretaceous of Europe and 

 the Middle East (Smith & Jeffery, in press). 



Material studied. MGB 37563, 37564, BMNH EE61 18-24. 



Order CALYCINA Gregory, 1900 



Diagnosis. Regular echinoids with stout elevated cap-like apical 

 system that incorporates one or more large suranal plates firmly 

 bound to the ring of genital plates. 



Family SALENIIDAE Agassiz, 1838 

 Genus SALENIA Gray, 1835 



Diagnosis. Periproct offset towards ocular I. 



Subgenus SALENIA (PLEUROSALENIA) Pomel, 1883 



Diagnosis. Salenia with predominantly simple ambulacral plat- 

 ing; plating entirely simple towards peristome. 



Salenia (Pleurosalenia) scabra (Nestler, 1965) 



PI. l,figs 13, 14; Fig. 6 



1 950 Salenidia bonissenti Cotteau; Kongiel: 3 1 1 , 32 1 , pi. 1 , figs 



1^. 

 1965 Salenidia scabra Nestler: 987, pi. 4, figs 4-7. 

 1973 Salenidia scabra Kutscher: ill, figs 13, 14. 

 1975 Salenidia scabra Nestler: 89. 

 1 979 Salenidia cf. bonissenti Cotteau; Geys: 890, pi. 1 , fig. 4, pi. 



2. fig. 2, pi. 3, figs 6-8. pi. 4, figs 1, 2. 

 1 987 Salenidia ?bonissenti Van der Ham et al.: 21. 



Diagnosis. Up to 15 mm diameter; relatively tall (height ca. 60% 

 test diameter). Apical disc subcorneal, weakly pentastellate in out- 

 line with ocular plates projecfing slightly. Periproct small, trigonal, 

 with slight lip. Large oval sutural pits at triple junctions and mid- 

 length along all disc plate sutures; strong ridges run radially between 

 the sutural pits giving the disc a very characteristic appearance. 

 Ambulacra straight, unigeminate throughout (rare bigeminate ele- 

 ments at ambitus), with relafively wide perradial band of granulation 

 separating primary tubercles. Five interambulacral plates in a column. 



Occurrence. Maastrichdan of Santander, horizon 4. Elsewhere 

 the species is known from the late Early Maastrichtian of Riigen, 

 Germany, Late Maastrichdan of Poland and Eben-Emael, Belgium. 



Material studied. 

 37575. 



BMNH EE4423a, b, BMNH EE4430, MGB 



Remarks. Separated from S. anthophora at comparable test diam- 

 eters by having a smaller sized disc. Salenia bonissenti (Cotteau, 



