LATE CRETACEOUS-EARLY TERTIARY ECHINOIDS 



93 



Fig. 6 Camera lucida drawings of plating in Salenia (Pleurosalenia) 

 scabra (Nestler, 1965) from the coast west of Cabo Mayor (Santander, 

 Cantabria); BMNH EE6I 10. a. apical disc; b, adapical ambulacrum; c, 

 adoral ambulacrum, peristome at base. Scale bar = 1 mm. 



1 866b) has usually been treated as a distinct species ciiaracterized by 

 having %harp caiina on apical disc plates (e.g. Geys, 1 979). However, 

 Cotteau's original illustrations do not show sharp carina, only a 

 somewhat gentle undulose plate surfaces similar to many specimens 

 of S. anthophora. The oldest available name for forms with sharp 

 carina radially arranged on genital and suranal plates is Salenidia 

 scabra Nestler (1965). 



Salenia (Pleurosalenia) maestrichtensis Schluter, 1 892 



PI. 1, figs 8-10 



1857 Salenia bourgeoisi Bosquet: 3. 



1862 Salenia bourgeoisi Cotteau, in Cotteau 1861a-67a: 162 



{pars), pi. 1040, figs 25-28. 

 1892 Salenia Maestrichtensis Schluter. 196. 

 1935 Salenia bourgeoisi Cotteau; Smiser: 26, pi. 2, fig. 7. 

 1979 Salenidia maestrichtensis Schluter; Geys: 298, figs 1.1- 



1.7. 

 1979 Salenidia sanctipetri Geys: 303. figs 5. 1-5.6, 6. 1-6.2. 

 1987 Salenidia maestrichtensis SchXulQV.'WdinAtv Warn etal.: 21, 



pi. 2, fig. 3. 

 1987 Salenidia sanctipetri Geys; Van der Ham et al.: 22. 



Diagnosis. Test generally rather small (up to about 12 mm test 

 diameter), and depressed in profile (height ca. 50% of test diameter). 

 Apical disc flat with raised margins and a small but distinct rim 

 surrounding the periproct. Periproct in general rather small, slightly 

 smaller than the suranal plate. Ocular plates T-shaped, projecting 

 beyond genital plates slightly. Sutural pits large, present at all triple 

 junctions and in addition midway along genital-ocular plate sutures; 

 becoming quite angular in large individuals. Ambulacra straight and 

 relatively wide, with outer columns of configuous primary tubercles 

 and an inner band of secondary tubercles with intervening granules. 

 Rare bigeminate elements may be present, but for the most part 

 ambulacra are unigeminate. Interambulacra wide with primary tu- 

 bercles posifioned close to the adradii, leaving a broad, somewhat 

 depressed interradial zone of secondary granules. Primary tubercles 

 with relafively coarse mamelons. Peristome small and somewhat 

 invaginated. 



Occurrence. Spanish material comes from the Maastrichtian, 

 horizon 6. Olazagutia Pass, Navarra province and Maastrichtian, 

 horizon 4. Santander. The species was first described from the Late 

 MaastrichUan Chalk of the Maastricht district. The Netherlands and 

 Belgium, and is also reported from the Ciply Craie phosphatee and 

 St Symphorien Gravel at Ciply, Belgium (fide Geys, 1979). 



Material studied. BMNH EE4423a, b, EE4430, MGB 37574. 



Remarks. The distinction between this species and juvenile 5. 

 anthophora is difficult. In both species apical disc plates are charac- 

 teristically dimpled at small sizes, and it may be that the two forms 

 cannot be separated until test sizes of about 1 cm have been reached. 

 By this size 5. maastrichtensis is obviously flatter and has wider and 

 more depressed interradial bands than S. anthophora. 



Subgenus SALENIA (SALENIA) Gray, 1835 



Diagnosis. Salenia with predominantly bigeminate ambulacral 

 plating; plating never simple adorally. 



Salenia (Salenia) sp. 



Diagnosis. A small individual 4.2 mm in diameter with a large 

 apical disc with prominent sutural pits. Ambulacral plating bigeminate 

 at ambitus and adorally. 



Material studied. BMNH EE443 1 . 



Occurrence. Maastrichtian, horizon 9, Santander. 



Remarks. This specimen is clearly distinct from the other saleniids 

 described here on account of its bigeminate ambulacral plating. 

 However, it is too small an individual to identify with confidence to 

 species level. 



Order ARBACIOIDA Gregory, 1900 

 Family GONIOPYGIDAE Smith & Wright, 1993 



Genus GOA'/O/'FGt/S Agassiz, 1838 



Diagnosis. Apical disc solid and raised above the corona; no 

 suranal plate present; genital plates pointed with gonopore opening 

 at tip or in immediately adjacent interambulacra. Primary tubercles 

 imperforate, non-crenulate. Ambulacral plating basically trigeminate 

 with upper element in each triad occluded. 



Goniopygus tetraphyma Lambeil, 1907 PI. 1, figs 16, 17 



1907 Goniopygus tetraphyma Lambert: 701, pi. 25, figs 7-9. 

 1949 Goniopygus jeanneti Sanchez Roig: 54, pi. 2. figs 7, 8. 



Diagnosis. Test up to 25 mm diameter; subconical in profile with 

 low ambitus and small flat summit. Apical disc solid, elevated above 

 the corona; dicyclic. with pointed genital plates; gonopores opening 

 at the very distal point of genital plates, beyond the elevated portion. 

 Periproct small, trigonal, with three perianal tubercles. Ambulacra 

 wide, trigeminate throughout with small phyllodes adorally; each 

 plate carrying a large primary tubercle and a smaller secondary 

 tubercle, the latter forming a distinct inner series. Interambulacra 

 with single large primary tubercle; adapical plates lacking secondary 

 tubercles, but those from the ambital region adorally have five or six 

 small secondary tubercles. Peristome large, hardly sunken. 



Occurrence. Maastrichtian, horizon 7/8. Santander. The type of 

 this species comes from the Upper Cretaceous of Roquefort (Haute- 



