94 



A.B. SMITH ETAL. 



Garonne, France). It is also known from the Upper Cretaceous of 

 Cuba and from the Maastrichtian of Fi'gols, Llobregat River basin, 

 Catalonia, Spain (MGB collections). 



Material studied. BMNH EE4419, EE6129, MGB 37576. 



Remarks. Easily distinguished from other species of Goniopygus 

 by its very wide ambulacra with prominent double columns of 

 secondary tubercles. Only the 25 mm diameter individual shows the 

 distinct double ambulacra! tubercles characteristic of this species. 

 Two very much smaller specimens (less than 10 mm) from the same 

 horizon are tentatively assigned to this species, although they are too 

 small to have developed prominent secondary ambulacral 

 tuberculation. 



oped. Distinct interradial naked zone developed adapically. 



Occurrence. Our two specimens come from the Upper Thanetian, 

 P. pseudomenardii Zone, limestone of Casas de Oraien. near Larumbe, 

 Navarra, Spain. The type locality for this species is the Thanetian of 

 Belbeze, Haute-Garonne, France. It has also been recorded from the 

 Thanetian of Le Tuco near Cazeneuve, Haute-Garonne, France, and 

 from the Petites Pyrenees (Plaziat et al. 1975). 



Material studied. BMNH EE 6117, MGB 37425. 



Remarks. The arcuate uniserial arrangement of ambulacral pore- 

 pairs distinguish this species from other phymosomatid species 

 described here. 



GQxm% DIPLOTAGMA Schluter, 1870 



Order PHYMOSOMATOIDA Mortensen, 1904 

 Family PHYMOSOMATIDAE Pomel, 1883 



Diagnosis. Apical disc not firmly bound to corona; large and 

 monocyclic, but plating usually lost. Ambulacra polygeminate with 

 phymosomatid-style compounding. Primary tubercles imperforate 

 and crenulate. 



Genus GAUTHIERIA Lambert, 1888 



Diagnosis. Phymosomatids with pore-pairs uniserial throughout. 

 One large interambulacral tubercle on ambital interambulacral plates 

 with small adradial tubercles present on adoral plates. Peristome 

 usually deeply sunken. 



Gauthieria pseudomagnifica (Cotteau, 



1877) 



PI. l,figs 18-20 



1863 Cyphosoma magnificum Cotteau: 185 {nomen nudum). 

 1 877 Cyphosoma pseudomagnificum Cotteau; Cotteau: 55, pi. 4, 

 figs 1-6. 



Diagnosis. Test up to 35 mm in diameter; circular in outline and 

 depressed in profile (height about 40-50% of diameter). Apical disc 

 opening pentagonal, relatively small (30% of test diameter). 

 Peristome equally small and deeply invaginated. Ambulacra taper- 

 ing adapically; ambital compound plate with six or seven pore-pairs 

 defining weak arcs; pore-pairs remain uniserial throughout, with no 

 pore-crowding adorally. Single large primary tubercle on each 

 ambulacral and interambulacral plate. Tubercles large with circular 

 areole and relatively small mamelons. Much smaller secondary 

 tubercles present on adradial and interradial margins of subambital 

 plates. Granulation outside primary tubercles rather poorly devel- 



Dl AGNOSIS . Phymosomatid with biserial pore-pairs developed from 

 the ambitus adapically and with strong adoral phyllodes. Test 

 subconical and apical disc rather small. Interambulacral plates very 

 wide and low with primary tubercles contiguous in column; broad 

 granular zones developed both interradially and perradially. Tuber- 

 cles imperforate and crenulate. 



Diplotagma sp. 



PI. 2, figs 1,2 



Diagnosis. Test 37-38 mm in diameter and about 23 mm in 

 height. Apical disc about 50% of test diameter, peristome slightly 

 larger and a little sunken. Pore-pairs arranged biserially from the 

 ambitus adapically but in arcs subambitally and forming phyllodes 

 adorally. Subambital plates have five pore-pairs to a plate, adapical 

 plates with four pore-pairs. Ambulacral tubercles are contiguous in 

 vertical columns and are separated by a wide perradial zone of 

 granulation. Interambulacral plates are very wide (more than three 

 times broader than tall at the ambitus) and have a single primary 

 tubercle. Secondary tubercles are developed adradially, one on oral 

 plates and up to three on ambital and supra-ambital plates. 



Occurrence. Maastrichtian, horizon 7/8, Santander. 



Material studied. BMNHEE4417. 



Remarks. Our only specimen is rather badly abraded, but shows 

 the general features of Diplotagma, namely the rather delicate 

 primary tubercles and broad granular adradial and interradial zones, 

 together with the biserially arranged pore-pairs from the ambitus to 

 the apex. Its strongly developed adoral phyllodes distinguishes it 

 from Acanthechinus. Diplotagma sp. nov. (Smith & Jeffery, in 

 press), from the Upper Maastrichtian of the Maastricht area, differs 

 from the Spanish specimen in having narrower, coarser and more 

 heterogeneous interradial tuberculation. 



PLATE 2 



Figs 1, 2 Diplotagma sp., BMNH EE44 1 7, Maastrichtian of Santander, Cantabria. Oral and aboral views, x 1 .5. 



Figs 3, 4 Phymosoma granulosum (Goldfuss, 1829), BMNH EE6I27, Maastrichtian of Santander, Cantabria. Apical and lateral views, x2. 



Figs 5-8 Adelopneustes emsti Smith & Gallemi, BMNH EE6134 (holotype), Thanetian of Casas de Oraien, Navarra. Apical, oral, lateral and posterior 



views, x2. 

 Figs 9-11 Camerogalerus cantabricus Smith & Gallemi. BMNH EE6132 (holotype), Maastrichtian of Santander, Cantabria. Oral, apical and lateral 



views, x5. 

 Fig. 12 Coenholectypus nachtigali (Krumbeck, 1906), BMNH EE6129, Maastrichtian of Santander. Cantabria. Apical view, xl .5. 

 Figs 13-16 'Echinogalerus' vetschauensis (Schluter, 1902), BMNH EE6164, Maastrichtian of Santander. Cantabria. Apical, oral, lateral and posterior 



views, x3. 

 Figs 17-20 Echinogalerus muelleri (Schluter, 1902), BMNH EE6138, Maastrichtian of Olagazutia Pass, Navarra. Apical, oral, lateral and posterior views, 



x3. 



