100 



A.B. SMITH ETAL. 



Fig. 9 Camera lucida drawings of plating in Adelopneustes emsti sp. nov. 

 from Casas de Oraien (Navarra); BMNH EE6134. a, apical surface; b. 

 oral surface; c, adapical ambulacrum; d, apical disc; e, subambital 

 ambulacrum. Scale bars; a, b = 2 mm; c-e = 1 mm. 



Genus ADfiLOPA^Et/Sr^SGauthier, 1889 



Diagnosis. Conulids with pyrinoid ambulacral plating orally and 

 simple plating aborally. Peristome subcircular to weakly elliptical. 



Adelopneustes emsti Smith & Gallemi, sp. nov. 



PI. 2, figs 5-8; Fig. 9 



Diagnosis. Test ovate in outline, slightly pointed to rear; 26 mm in 

 length, slightly narrower than long; low-domal in profile, with test 

 height 14.5 mm. Apical disc tetrabasal with large gonopores sur- 

 rounded by rims (female character only?); subcentral on genital 

 plates. Ambulacral pores very small and rudimentary, almost impos- 

 sible to see even under magnification; double adapically. Ambulacral 

 plating apparently simple aborally; plating adorally not visible. 

 Peristome oblique (crushed in the only specimen known), elongate 

 towards anterior left. Lower surface strongly rounded. Periproct 

 elongate and pointed both adorally and adapically, visible from 

 beneath and posteriorly; as large as mouth. 



Occurrence. Upper Thanetian, P. pseudomenardii Zone, Casas 

 de Oraien, Larumbe, Navarra province, Spain. 



Types. Holotype, BMNH EE6 1 34. 



Remarks. Differs from A', boehmi (Neitsch) in having a more 

 depressed profile and larger gonopores surrounded by a raised 

 platform. Differs from N. montainvillensis (Sorignet) in having an 

 oblique rather than circular peristome. Differs from both these 

 species and from all others assigned to this genus in having rudimen- 

 tary aboral pores. 



Order CASSIDULOIDA Claus. 1880 



Diagnosis. Peristome small, lacking buccal notches; 

 interambulacra undifferentiated. Ambulacral pores differentiated 

 into aboral petals and oral phyllodes, though occasionally rudimen- 

 tary throughout. 



Family PLACHIOCHASMIDAE Smith & Jeffery, in press 

 Genus ECHINOGALERUS Konig, 1825 



Diagnosis. Peristome small, anterior of centre. Periproct slightly 

 wider than long, inframarginal. Petals and phyllodes rudimentary. 



Echinogalerus muelleri (Schliiter, 1902) 



PI. 2, figs 17-20; Fig. 10b, d 



1902 Caratomus Muelleri Schluter: 316, pi. 1 1, figs 14-17. 



1927 Echinogalerus belgicus Lamhen: 38. 



1987 Echinocyamus muelleri SchlnteT^VandeTHam et al.: 28, pi. 



9. fig. 3.' 

 1992 Echinogalerus muelleri Schluter; Van der Ham & Van 



Birgelen: 149, pi. 3. fig. 7. 

 1992 Echinogalerus belgicus GaWemi: 170, photo 6. 



Diagnosis. Test up to 15 mm in length; width about 85-90% of 

 length; height about 60% of test length. Ovate in outline with 

 rounded anterior and widest point slightly posterior to midlength. 

 Depressed in profile with rounded margins; lower surface strongly 

 pulvinate and slightly sunken towards peristome; upper surface 

 depressed. Ambulacra composed entirely of simple plating; 

 ambulacral pores double throughout, almost subpetaloid adapically 

 in largest individuals; uniserial adorally. Pore-pairs oblique and 

 strictly uniserial towards the peristome; widely separated. Peristome 

 slightly anterior of centre; relatively large (13% of test length); 

 oblique along 3-V axis. Periproct subambital, clearly visible in oral 

 view; transverse, wider than the peristome, subtrigonal and pointed 

 adorally; separated from the peristome by about 4 interambulacral 

 plates. Tuberculation of scattered sunken tubercles. 



Occurrence. Maastrichtian, horizons 3 to 6, Santander, Cantabria; 

 Maastrichtian, horizon 6, Olazagutia Pass, Navarra province. Lower 

 Maastrichtian, Homes Morts Member, Salas de Pallars and Sapeira, 

 Tremp Basin (Gallemi 1992). The type comes ft'om the Upper 

 Maastrichtian of Vetschau, near Aken, Germany. 



Material STUDIED. BMNH EE4412(a-d), EE4413(a-f), EE6137- 

 38, EE6163; MGB 37451-57, 37490-98, 37500-02, 37526, 37528, 

 37532, 37537, 37546. 



Remarks. Similar in shape to E. belgicus. but with double pores in 

 its ambulacra. Differs from E? vetschaupensis in having a larger 

 peristome, more pulvinate lower surface and strictly uniserial pore- 

 pairs adorally. Populations from Tremp and from Olazagutia Pass 

 contain both large and small individuals, and are assigned to this 

 species without doubt. However, only small individuals have so far 

 been found at horizons 3 and 5 at Santander. Because these are 

 indistinguishable from small individuals that we have from 

 Olazagutia, we have included them in this species. 



Echinogalerus? vetschauensis (Schliiter, 1902) 



PI. 2, figs 13-16; Fig. 10a, c 



1902 Caratomus vetschauensis Schluter: 318. pi. 11, figs 10-13. 

 1 987 Echinogalerus vetschauensis Schluter; Van der Ham et al. : 

 28, pi. 9, fig. 5. 



