102 



A.B. SMITH ETAL 



Fig. 11 Camera lucida drawing of plating in Clypeolampas ovatus 

 (Lamarck, 1816) from the Maastrichtian of the coast west of Cabo 

 Mayor (Santander, Cantabria); BMNH EE6144, phyllode of 

 ambulacrum II, peristome margin at base. Scale bar = 1 mm. 



Diagnosis. As for genus. 



Occurrence. Maastrichtian, horizons I, 2 and 4, Santander; 

 Maastrichtian, Somo Beach, near Santander, Cantabria province. 

 Upper Campanian of Barranc d" en Jaume, Quatretonda and Nicolasa 

 quarry, Tavemes, Valencia province, Spain (Nickles 1892). This 

 species is widely reported from the Maastrichtian of the Anglo-Paris 

 Basin, Charente-Maritime and Dordogne, France. 



Material studied. BMNH EE6 1 44-^7 . 



Remarks. This species is moderately common in the shallowest 

 limestones at the base of the transgressive Maastrichtian sequence at 

 Santander Several species have been distinguished by Lambert 

 (1920) on material from the Santander region. The differences 

 between his species C. mengaudi, C. douvillei and C. ovatus are 

 based on the disposition of pores in the phyllodes. Although there are 

 certainly slight differences, it is not at all clear how much individual 

 variation exists in this character within single populations, and the 

 three taxa may well turn out to be conspecific. 



Unnamed Family 

 Genus Aff/CLEO/'FCl/S Agassiz, 1840 



reported here, by its large size, strongly transverse peristome and 

 posterior longitudinal periproct. There are many species of 

 Gitolampas recognized from the Maastrichtian and Palaeocene 

 (Smith & Jeffery, in press). It is distinguished from the closely 

 similar G. ataxensis (Cotteau) by the form of its petals, which are 

 more bowed and proportionally longer than in G. ataxensis. 



Diagnosis. Subquadrate in outline and depressed in profile, with 

 supramarginal periproct opening into subanal sulcus. Apical disc 

 tetrabasal. Ambulacra subpetaloid at most. Peristome lacking 

 bourrelets; perioral tuberculation confined to peristomial well. 

 Phyllodes generally rather undeveloped with every third pore-pair 

 insert. 



Family CLYPEOLAMPADIDAE Kier, 1962 

 Genus CLYPEOLAMPAS Pomel, 1869 



Diagnosis. Test with flat base and highly vaulted upper surface. 

 Petals long, extending almost to the ambitus; slightly expanding and 

 widely open distally. Periproct submarginal at rear of oral surface; 

 transverse. Narrow granular sternal zone present behind periproct. 

 Phyllodes widely expanded, with many pores scattered between the 

 inner and outer series. Interambulacra swollen towards peristome 

 with 'bourrelets' composed of many small plates rather than just a 

 single large basicoronal plate. 



Clypeolampas ovatus (Lamarck, 1816) 



PI. 3, figs 3-5; Fig. 11 



1816 Galerites ovatus Lamarck, 1816: 22. 



1829 Clypeaster leskei Goldfuss. 1829: 132, pi. 42, fig. 1. 



1920 Clypeolampas leskei Goldfuss; Lambert: 21. 



1996 Clypeolampas cf. ovatus (Lamarck); Wilmsen et ai: fig. 7. 



Nucleopygus coravium (Agassiz, in Agassiz & Desor, 

 1847) PI. 3, figs 6-9; PI. 4, figs 7-9 



1 847 Nucleopygus cor avium Agassiz; Agassiz & Desor: 1 52. 



1898b Nucleopygus coravium Lambert: 168. pi. 5, figs 1^, 8-14. 



1935 Nucleopygus coravium Smiser: 52, pi. 5, fig. 1. 



1987 Nucleopygus coraviumVandeTliametal. -.29,^1. 11, fig. 2. 



Diagnosis. Small species, rarely reaching more than 10 mm in 

 length; width ca. 75% of length; depressed with height ca. 50% of 

 length. Ovoid in outline with a rounded anterior and slightly trun- 

 cated posterior; widest point posterior of centre. In posterior profile 

 the test is strongly concavo-convex. Apical system anterior, about 

 one-third of test length from the anterior; tetrabasal with posterior 

 genital plates not separated by the madreporite; genital plate 3 

 smaller than the remainder Petals small and rudimentary, particu- 

 larly the anterior petal ; lateral and posterior petals equally developed, 

 the posterior pair ending well before reaching the level of the 

 periproctal opening. Pore-pairs weakly conjugate with equal-sized 

 pores. Peristome large, oval to subpentagonal, slightly anterior of 

 centre, depressed with inwardly sloping margins. Phyllodes more or 



PLATE 4 



Figs 1-6 Nucleopygus scrobiculatus (Goldfuss, 1829), Maastrichtian of Santander, Cantabria. 1, BMNH EE6157; apical view, x2. 2-4, BMNH 6156; oral, 



lateral and posterior views, x2. 5, 6, BMNH EE6154; oral and posterior views, x2. 

 Figs 7-9 Nucleopygus coravium (Agassiz, in Agassiz & Desor, 1 847), BMNH EE4403, Maastrichtian of Olazagutia Pass, Navarra. Apical, lateral and oral 



views, x3. 

 Figs 10-13 Offaster leymeriei Cotteau, 1887, BMNH EE6170, Maastrichtian of Santander, Cantabria. Apical, oral, lateral and posterior views, xl.5. 

 Figs 14-18 Galeaster bertrandi Seunes, 1889, BMNH EE6188, Maastrichtian of Santander, Cantabria. Apical, oral, posterior, anterior and lateral views, 



x2. 

 Fig. 19 Cardiasler ip. BMNH EE6174, Thanetian of Casas de Oraien, Navarra. Oral view, xl. 

 Figs 20-22 Echinocorys scutata (oTma. cotteaui Lambstt, 1903, BMNH EE6 189, Thanetian of Casas de Oraien, Navarra. Oral, lateral and apical views, xl. 



J 



