LATE CRETACEOUS-EARLY TERTIARY ECHINOIDS 



a_ >-. b 



105 



Fig. 13 Camera lucida drawings of plating in Oolopygus sp. and Zuffardia sp. from the Maastrichtian of the coast west of Cabo Mayor (Santander, 

 Cantabria); a. Oolopygus sp., BMNH EE6169, apical disc; b. c, Zuffardia sp., BMNH EE6153; b, apical disc; c, phyllode and peristome margin. Scale 

 bars = 1 mm. 



and with V-shaped phyllodes comprising a single series of pores. 

 Periproct supra-ambital, on the underside of a large lobate projec- 

 tion; opening appears wide and slit-like in posterior profile. Oral 

 tuberculation arranged with a strong bilateral symmetry. 



Rhynchopygus sp. 



Occurrence. Maastrichtian, horizon 4, Santander 



Material studied. BMNH EE6162. 



Remarks. Our record is based on a single specimen, representing 

 the posterior portion of a test. This shows the highly characteristic 

 lobe-like projection above the wide and slit-hke periproctal opening. 

 There are two species of Rhynchopygus, R. mannini {Agassiz) and R. 

 donetzensis (Faas), differing primarily in the development of their 

 petals. As our specimen does not show the petals, it cannot be 

 assigned to species level. 



Genus RHYNCHOLAMPAS Agassiz, 1 869 



Diagnosis. Small cassiduloids with a flat base and ovate outline. 

 Peristome small, subcentral, weakly transverse to as wide as long; 

 rim of peristome hardly swollen; deep well leading into peristomial 

 opening. Periproct supra-ambital; transverse, hardly sunken. 



Rhyncholampas macari (Smiser, 1935) PI. 3, figs 15. 16 



1935 Rhynchopygus macari Smiser: 63, pi. 6, fig. 6. 

 1987 Procassidulus macari (Smiser); Van der Ham et a!.: 30, pi. 

 11, fig. 6. 



Diagnosis. Test up to 33 mm in length; elongate with flat base and 

 depressed upper surface. Width 75-80% of length, widest posterior 

 of mid-length; height no more than 35% of length. Apical disc a little 

 anterior of centre, with four gonopores; probably tetrabasal, but 

 plating not seen. Petals well-developed, bowed and converging 

 distally; posterior petals shorter than lateral and anterior petals. 

 Peristome slightly anterior of centre; small and pentagonal; almost 

 as wide as long; surrounded by small but distinctly swollen bourrelets. 

 Phyllodes composed of a V-shaped outer series of pores plus one or 

 two inner pores near the apex of the V. Broad pitted sagittal zone free 

 of tubercles both anterior and posterior to the peristome. Periproct 

 supramarginal; transverse, with slight aboral canopy. 



Occurrence. Maastrichtian, bed 4, Santander. This species also 

 occurs in the Lichtenberg Horizon, and Nekum and Meerssen Mem- 



bers, Maastricht Formation, B. junior and B. kazimiroviensis Zones, 

 Upper Maastrichtian of the Maastricht district. The Netherlands and 

 Belgium (Van der Ham et al., 1987). 



Material studied. BMNH EE6 150-52, MGB 37512 



Remarks. Although our material is unfortunately rather crushed, 

 all relevant plating details are seen, allowing us to be confident in our 

 determination. No other cassiduloid described herein has a trans- 

 verse periproct situated supra-ambitally. 



Family FAUJASIIDAE Lambert, 1905 

 Genus ZUFFARDIA Checchia-Rispoli. 1917 



Diagnosis. Globose cassiduloids with monobasal apical disc and 

 relatively short petals. Peristome small, subcentral, pentagonal, sur- 

 rounded by distinct phyllodes and bourrelets. Periproct posterior, 

 hardly invaginated. All pores below petals single. 



Zuffardia sp. 



PI. 3, figs 10, 11; Fig. 13b, c 



Diagnosis. Test up to 25 mm in length; almost subcircular in 

 outline, inflated in profile with flat to weakly convex base and 

 subcentral peristome. Apical disc compact, with four gonopores. 

 Petals relafively short and weakly bowed; extending about half the 

 distance to the ambitus. Peristome small, pentagonal, surrounded by 

 small bourrelets. There is a deep well leading into the peristomial 

 opening. A broad naked pitted zone is developed both anterior and 

 posterior to the peristome on the oral surface. Periproct posterior, 

 subcircular, hardly invaginated. Phyllodes well developed with bowed 

 outer series of ten pores separated from three inner series pores. All 

 pores single below the petals. 



Occurrence. Maastricthian, horizon 4, Santander. 



Material studied. BMNH EE6 1 53, MGB 3752 1 , 37525. 37527. 



Remarks. Resembles Catopygus fenestratus Agassiz & Desor in 

 general appearance, but distinguished from that species by having 

 entirely single pores below the petals, and in having a well-devel- 

 oped anterior and posterior naked zone that is pitted. This latter 

 feature distinguishes it from the type species Z. morgani (Cotteau & 

 Gauthier), which has an unpitted naked zone adorally. Very similar 

 material was described from the Late Maastrichtian, B. 

 kasimiroviensis Zone of Mangyshlak by Jeffery (1997). Our sped- 



