108 



A.B. SMITH ETAL. 



1907 Hemipneustes leymeriei Hebert; Lambert: 709. 



1935 Spatangoidesstria1oradiatus\^tskt\S>rmicx.l\,^\.^Si%- 1- 



1973 Hemipneustes leymeriei W&otx\.;K?Ld\g: 13. 



1987 Hemipnesutes striatoradiatus Leske; Van der Ham et al:. 



32, pi. 18, fig. 2. 

 1995 Hemipneustes striatoradiatus Gallemi et al.: table 1. 



Diagnosis. Test 66 mm in length; tall (height 75% of length); oval 

 in outline and only slightly longer than wide. Upper surface strongly 

 domed in both lateral and posterior profiles. Anterior sulcus narrow 

 and sharply defined at the ambitus. 



Occurrence. Maastrichtian, horizon 4, Olazagutia Pass, Navarra 

 province. ?Maastrichtian, horizon 2, Santander. 



Material studied. BMNH EE6179, MGB 37442-43. 



Remarks. Only three specimens of this species were found along 

 with the very much more common H. pyrenaicus Hebert at Olazagutia 

 Pass. They are easily distinguished from that species by their domed 

 profile and very much more circular outline. 



Fragments of a rather tall, domal Hemipneustes are found in the 

 lower beds at Santander. Unfortunately, these are too incomplete to 

 be certain of their specific idenfity. 



Hemipneustes pyrenaicus Hebert, 1875 PI. 7, figs 1, 2 



1875 Hemipneustes pyrenaicus W&otri: 593,^1. 19. 



1892 Hemipneusts arnaudi Cotleau: 165, pi. 21, figs 3, 4. 



1892 Hemipneustes pyrenaicus Hehert^Nickles: 110. 



1907 Hemipneustes pyrenaicus Hebert; Lambert: 709. 



1920 Hemipneustes pyrenaicus Hebert; Lambert: 25. 



1973 Hemipneustes pyrenaicus Hebert; Radig: 57, pi. 8, figs 6-9, 



pi. 9, figs 1^. 



1992 Hemipneustes pyrenaicus Hebert; Gallemi: 244, photo 13. 



71996 Hemipneustes pyrenaicus Hebert; Wilmsen et al.: 354. 



Diagnosis. Test up to 95 mm in length; elongate, with width 75- 

 80% of length; height 60-65% of length; subquadrate in profile with 

 slight peak centrally. Ambulacral groove narrow; apical disc central 

 to a little anterior. Periproct opening on inward-sloping portion of 

 test and thus just visible in oral view. 



Occurrence. Maastrichtian, beds 2-4, Olazagutia Pass, Navarra 

 province; Maastrichtian, Somo Beach near Santander, Cantabria 

 province; Lower Maastrichfian, Homes Morts Member, Salas de 

 Pallars, Tremp Basin, Catalonia (Gallemi 1977). 



Material studied. BMNH EE4381, EE6 180-87, MGB 37472, 

 37474-75, 37477. 



Remarks. Readily distinguished from H. striatoradiatus by its 

 more elongate form, more inwardly sloping posterior face and more 

 depressed profile. 



Hemipneustes sp. 



1992 Hemipneustes leymerei Gallemi: 240, photo 12. 



Diagnosis. Test longer than 80 mm (damaged at rear, but prob- 

 ably approximately 90 mm); rounded, almost circular with width 

 very probably representing 90-95% of length; height around 55% of 

 length; subrectangular in profile with a slight depression centrally 

 where the apical system lies. Ambulacral groove narrow at the 

 ambitus and expanding moderately towards the frontal part of the 

 apical surface. Periproct unknown. 



Occurrence. Maastrichtian, horizon 4, Santander. Homes Morts 

 Member, Lower Maastrichtian, Salas de Pallars, Tremp Basin 

 (Gallemi, 1992). 



Material studied. MGB 37533. 



Remarks. Distinguishedfromf/./?vre«a/cM5and//. striatoradiatus 

 by its rounded, almost circular form and from the latter species also 

 by its more depressed, flat-topped profile. 



Family ECHINOCORIDAE Lambert, 1917 

 Germs ECHINOCORYSh&sk&, 1778 



Diagnosis. Test hemispherical to subconical with flat base; no 

 frontal groove. Apical disc central; holasterid in form with four 

 gonopores; madrepores confined to genital plate 2. Peristome D- 

 shaped and downward facing; test generally slightly depressed 

 towards opening. All ambulacra similar, non-petaloid; pore-pairs 

 small and double aborally. Periproct transverse; on lower surface. 

 Plastron meridostemous. No fascioles. 



Remarks. Echinocorys has been divided into a large number of 

 species on the basis of overall shape. Although individual 

 morphologies can be very distinctive and of great use for local 

 correlation, there is also a great deal of integradation amongst forms, 

 with similar extremes of shape recurring at different times in the 

 history of the group. As there are no structural differences in the form 

 of the test, we here assign all forms to one large species complex, E. 

 scutata Leske, and recognize the various shape varieties as named 

 forms. 



Echinocorys scutata Leske, 1778 



1778 Echinocorys scutata Leske: 1 1 1, pi. 15, figs a, b. 



Diagnosis. Ambulacral plates becoming low and densely packed 

 adapically; aboral pore-pairs small and circumflexed, becoming 

 denser adapically, but not conjugate and not forming distinct petals. 



Echinocorys scutata forma ovata Leske, 1778 



Fig. 17a 



1778 Echinocorytes ovatus Leske: 178, pi. 53, fig. 3. 

 1801 Anachytes ovatus Lamarck: 347. 



1903 Echinocorys ovatus Leske; Lambert: 69, pi. 4, figs 6, 7, pi. 

 5, figs 1,2. 



Diagnosis. Test up to 90 mm in length and ovate in outline; width 

 approximately 75-85% of length. Subconical in profile, varying 

 from flat-topped to distinctly pointed. Height 67-80% of length. 

 Ambitus low and a little depressed towards the peristome. Aboral 

 pore-pairs small and circumflexed; becoming more crowded 

 adapically, but not forming distinct petals and never conjugate. 



Occurrence. Maastrichtian, horizon 3, Santander, Cantabria; 

 Lower Maastrichtian, black-shale facies, Sarasate, motorway sec- 

 tion, Navarra. 



Material studied. BMNH EE4567, BMUW 94788, 95005, 

 95006, 95014, 94780, 94787. 94786, 95012-13, 94783, 94788, 

 94996-98,94791. 



Echinocorys scutata forma belgica Lambert, 1 898 



Fig. 17b 



1829 Anachytes sulcatus Goldfuss: 146 (pars), pi. 45, fig. la-c 



only. 

 1898b Echinocorys belgicus Lambert: 43, pi. 4, figs 9, 10. 

 1 903 Echinocorys belgicus Lambert; Lambert: 80. 

 1996 Echinocorys belgicus Lambert; Wilmsen et al.: 354. 



