LATE CRETACEOUS-EARLY TERTIARY ECHINOIDS 



109 



Fig. 17 Outlines of tests of Echinocorys scutata Leske in lateral profile 

 (anterior to the right), a, forma ovata Leske, 1778. BMNH EE6194. 

 Maastrichtian of Santander, Cantabria. b. forma belgica Lambert, 1898, 

 BMNH EE6193, Maastrichtian of Santander. c, forma cotteaui Lambert, 

 1903, BMNH EE6191, Late Thanetian of Casas de Oraien, Navarra. d, 

 ionnsipyrenaica Seunes, 1888, BMNH EE6192, Danian of Erica, 

 Navarra. 



Echinocorys scutata forma cotteaui Lambert, 1903 



PI. 4, figs 20-22, PI. 7, figs 5, 6; Fig. 17c 



1847 Aiumchyies semiglobus Goldfuss: Agassiz & Desor: 136 



(pars). 

 1856 Echiiiocorys vulgaris Breynius; Leymerie & Cotteau: 343. 

 1877 Echinocorys semiglobus Goldfuss; Cotteau: 62, pi. 5, figs 



14-18. 

 1 887b Echinocorys semiglobus Goldfuss; Cotteau: 662. 

 1890 Echinocorys semiglobus Go\di\x's'i.S&une.s: 194. 

 1903 Echinocorys cotteaui Lambert: 84, pi. 4. fig. 8. 

 1907 Echinocorys cotteaui Lambert; Lambert: 7 1 8. 



Diagnosis. Test small, up to 70 mm length only; width 85-95% 

 test length, ovate in outline. Test low conical in profile, height 65- 

 73% of test length. Ambulacra simple and identical, with pore-pairs 

 becoming moderately dense towards the apex, but remaining small 

 and semi-colon-like; no clearly defined petaloid area. Peristome 

 slightly wider and perioral area more depressed, forming character- 

 istic rounded anterior rim. 



Occurrence. Upper Thanetian. Casas de Oraien. Navarra prov- 

 ince, Spain. 



Material studied. BMNH EE4530-4534. EE6 189-91, MGB 



37403-37405, 37429-37432. 



Diagnosis. Test 62 mm in length and 58 mm in width (width 94% 

 of length); ovate, with rounded anterior and slightly pointed poste- 

 rior. Height 35 mm (56% of length); with broad, rather flat, upper 

 surface, steeply sloping anterior and more gently sloping posterior 

 Ambitus very low. Lower surface distinctly sunken inside margins. 

 Petals small and non-conjugate; becoming denser towards apex. 



Occurrence. Maastrichtian, level 5, Santander 



Material STUDIED. BMNHEE6194. 



Remarks. Distinguished from other forms by its wide, subcircular 

 outline and depressed lateral profile. 



Echinocorys scutata forma pyrenaica Seunes, li 



Fig. 17d 



F 



I 



Echinocorys pyrenaicus Seunes; 814, pi. 30, fig. 5, pi. 31, 

 fig. 2. 

 1927 Echinocorys obliquus [Nilsson MS] Ravn: 336, pi. 4, fig. 2, 

 pi. 5, fig. 2. 



Diagnosis. Small form up to 50 mm in length; width 85-90% of 

 length; ovate and slightly pointed posteriorly. Flat base with sharp 

 and low ambitus. Upper surface conical in lateral and posterior 

 profile; height approximately 50% of test length. Periproct on oral 

 surface. Aboral pore-pairs small; becoming more closely-spaced 

 adapically. 



Occurrence. Danian Corajrer Beds, Erice, Navarra. Also known 

 from the Danian of Alfas del Pi, Alicante, Spain. The type of obliquus 

 comes from the Upper Danian of Denmark. 



Material studied. BMNH EE6192, MGB 37323. 



Remarks. The strongly conical profile and sharp ambitus disUn- 

 guish this form from all others. 



Remarks. Differs from forma belgica in profile; that form being 

 very much more flat topped. Less conical and with a more rounded 

 ambitus than forma, pyrenaica. 



Gems JERONIA Seunes, 1888 



Diagnosis. Ovate in outline with domed upper surface and flat 

 base; distinctly pointed at posterior Apical disc holasterid, with 

 three gonopores (genital plate 2 without a gonopore). Aboral pore- 

 pairs small and insignificant, placed centrally towards lower edge of 

 ambulacral plates and not crowded or enlarged adapically. Peristome 

 ovate and downward-facing; rather strongly depressed; oval. Periproct 

 inframarginal, opening on posterior projection. Tuberculation gen- 

 erally fine, but with prominent band of very large primary tubercles 

 around the ambitus. Plastron plating meridostemous; very narrow. 



Remarks. Similar to Echinocorys in shape, but with a stronger 

 sternal keel and distinct posterior point, and with a snpra-ambital 

 band of large tubercles around the posterior of the test. 



Jeronia pyrenaica Seunes. 1 i 



Figs 18-20 



1888a Jeronia pyrenaica Seunes: 810, text-figs 1-3, pi. 30, fig. 1. 



71889 Ojfaster cuneatus Seunes: 806, pi. 24. fig. 3. 



1891 Jeronia pyrenaica Seunes: 29, text-figs 4. 5, pi. 3, figs 1,2. 



1975 Jeronia pyrenaica S>tunts\Y\diZ\a.X et al.:6?<A.f\o. 10. 



Diagnosis. As for genus. In both specimens which show the 

 apical disc plating genital plate 2 has only madrepores and no 

 gonopore, as first illustrated by Seunes (1891). 



Occurrence. Near base ofchalkfacies; Upper Danian, Larumbe; 

 basal Thanetian, G. angulata Zone (ZB 5). Aristregui. Navarra 

 province. 



MATERIAL studied. BMNH EE6 1 95. MGB 37353, 37382-37383. 



