LATE CRETACEOUS-EARLY TERTIARY ECHINOIDS 



119 



Fig. 28 Camera lucida drawings of plating in Galeaster bertmndi Seunes. 1 889, from the Maastrichtian of the coast west of Cabo Mayor (Santander, 

 Cantabria); BMNH EE6188, a, oral surface, interambulacra shaded; b, apical surface. Scale bar= 1 m^m. 



Genus M/C/L45r£« Agassiz, 1836 

 Subgenus /SOM/C/MSrS/? Lambert, 1901 



Diagnosis. Test cordate, with shallow anterior sulcus; subconical 

 in profile with periproct very low on short posterior face. Apical disc 

 ethmophract with four gonopores. Anterior ambulacrum sunken 

 with pore-pairs similar to those of other ambulacra. Peristome 

 forward-facing and placed almost at the anterior border, largely 

 hidden in oral view by the projecting labral plate. No subanal 

 fasciole. 



Remarks. The forward facing peristome and strongly labiate 

 labral plate ally this lineage with the type species, M. coranguinum 

 Leske. It is distinguished, however, by its lack of a subanal fasciole 

 and by the highly conical profile of the test. 



Micraster (Isomicraster) stolleyi Lambeit, 1901 



1869 Epiaster gibbus Schliiter: 237, pi. 2, figs 1, la-c. 



1901 Isomicraster stolleyi Lambert, in Lambert 1895a-1901a: 



959. 

 1927 Isomicraster dallonii Lambert; Lambert: 48. 

 1975 Micraster stolleyi Lambert; Stokes: 79, fig. 30f. 

 1989 Micraster (Isomicraster) aff. stolleyi Lambert; Kiichler & 



Kutz: 196, text-fig. 2. 

 1 992 Micraster stolleyi Lambert; Gallemi: 335, figs 88-90. 



Diagnosis. As for genus. Test approximately 70 mm in length and 

 45 mm in height. 



Occurrence. ?Uppermost Campanian-?Lower Maastrichfian; 

 Erice, Navarra, Spain. 



Material studied. MGB 37304. 



Remarks. This unique specimen is crushed, but cannot be mis- 

 taken for any other species. It comes from the upper beds in the 

 shale-limestone succession immediately underlying the Danian 

 Coraster Limestones at Erice. This has been determined as Upper 

 Campanian - lowermost Maastrichtian (Kiichler & Kutz, 1989). 



Subgenus PSEUDOGIBBASTER Moskvin, 1983 



Diagnosis. Test cordate, with shallow anterior sulcus; inflated, 

 with vertically truncated posterior. Apical disc ethmophract with 

 four gonopores. Anterior ambulacrum sunken with pore-pairs differ- 

 enfiated from those of other ambulacra. Peristome almost non-labiate 

 and downward-facing. Subanal fasciole present. 



Remarks. The non-labiate peristome differentiates this from the 

 type species, M. coranguinum Leske, and from all Upper 

 Maastrichtian species of M/cra^/er from north-western Europe. It is 

 distinguished from Isaster and Cyclaster in having four gonopores 

 rather than three. 



Micraster (Pseudogibbaster) tercensis Cotteau, 1 863 



PI. 8, figs 8-12; Figs 29, 30 



1 856 Micraster brevis (Desor); Leymerie & Cotteau: 346. 



1863b Micraster cortestudinarium Goldfuss; Cotteau: 220 (55). 



1863b Micraster tercensis Cotteau: 221 (56). 



1 877 Micraster tercensis Cotteau; Cotteau: 69, pi. 7, figs 29-32. 



1975 Micraster tercensis Cotteau; Plaziat et al.: 640, fig. 10. 



Diagnosis. Test up to 65 mm in length, with distinct anterior 

 groove; tapering to a blunt point posteriorly. Width 100-1 10% of 

 length. In profile, subconical with rounded base and approximately 

 vertical posterior truncation. Apex subcentral to slightly anterior; 

 generally a little sunken. Petals moderately long and straight; open- 

 ended distally ; anterior pair extending 70% of the radial distance to 

 the margin, posterior pair extending only 55-65% of the distance. 

 Petals depressed with both pores slightly elongate. Anterior 

 ambulacrum with enlarged pores adapically (18-22 enlarged pore- 

 pairs), forming a shallow groove to the peristome. Labral plate 

 moderately long and narrow; sternal plates unequal, with left sternal 

 plate having a smaller contact surface with the labral plate than the 

 right sternal plate. Periproct small, rounded, positioned high on the 

 posterior face. No peripetalous fasciole. Aboral tuberculation con- 

 sisting of a scattering of larger primary tubercles set amongst a dense 

 uniform groundmass of miliaries. Subanal fasciole present, enclos- 

 ing four or five enlarged subanal pore-pairs. 



