126 



A.B. SMITH ETAL. 



Occurrence. Maastrichtian, Santander, Cantabria: horizons 2 

 (21 specimens), 3 (11 specimens), 4 (13 specimens), 5 (13 speci- 

 mens), 6 (6 specimens). Maastrichtian, horizons 4 and 5, Olazagutia 

 Pass, Navarra. Lower Maastrichtian, Homes Morts Member, Salas 

 de Pallars and Sapeira, Tremp area (Gallemi, 1992). Elsewhere this 

 species is known from the Upper Gulpen Formation and Lower and 

 Upper Maastricht Formation. Maastrichtian of the Maastricht dis- 

 trict. The Netheriands and Belgium (Van der Ham et ai, 1987), and 

 the Maastrichtian of the Mons Basin, Belgium (Smiser, 1935). 



Remarks. Many of the specimens are badly deformed and dis- 

 torted, making certain identification difficult. 



Hemiaster koninckanus d'Orbigny, 1855 



PI. 9, figs 1-7; Fig. 35a, d 



1855 Hemiaster koninckanus d'Orbigny: 250, pi. 885. 



1861 Hemiaster angustipneustes Cotleau &Tnger. 318,434, pi. 



53, figs 5-11. 

 1984 Hemiasterkoninkanusd'OrbignyyanderUam: 169,fig. 1. 

 1987 Hemiaster koninckanus d'Orbigny, Van der Ham et ai: 33, 



pi. 20, fig. 2. 



Diagnosis. Like H. prunella, but differing in the following ways: 

 (i) test more elongate, with width 75-85% of test length, rather than 

 almost as wide as long; (ii) in profile, the posterior face is undercut 

 and slopes inwards slightly and the tallest point lies posterior of the 

 apical disc; (iii) the anterior petals are less divergent in smaller 

 individuals, forming an angle of ca. 90-100°; (iv) the labral plate 

 projects slightly more over the peristome. 



Material studied. BMNH E6238-39, EE6250-5 1 . 



Occurrence. Maastrichtian, Santander: horizon 2 (5 specimens), 

 horizon 6 ( 1 specimen). Maastrichfian, horizon 2, Olazagutia Pass, 

 Navarra province. Elsewhere the species is recorded from the Upper 

 Gulpen Formation and Lower and Upper Maastricht Formation, 

 Maastrichtian of the Maastricht district. The Netherlands and Bel- 

 gium. 



Hemiaster Stella (Morton, 1830) 



PI. 9, figs 8-10 



1830 Spatangus Stella Morton: 245, pi. 3, fig. 1 1 . 



1877 Hemiaster nasutulus Sorignet; Cotteau: 65, pi. 6, figs 19, 



20. 

 1907 Hemiaster punctatus var. garumnica Lambert: 720 [nomen 



nudum]. 

 1907 Hemiaster punctatus var. arizensis Lambert: 721 [nomen 



nudum]. 

 1975 Hemiaster garumnicus Plaziat et ai: 643, fig. 10. 



Diagnosis. Globose test with convex anterior margin; test slightly 

 longer than wide. Apical disc set a little behind mid-length. Anterior 

 ambulacrum with long parallel-sided frontal groove adapically. An- 



terior petals slightly flexuous, diverging at about 80-90°; shorter 

 than anterior sulcus, but about twice as long as posterior petals. 

 Ovate peripetalous fasciole without indentations. 



Occurrence. Upper Thanetian, P. pseudomenardii Zone, Casas 

 de Oraien, Navarra province, Spain. Also known from the Thanetian 

 of Landes and the Petites Pyrenees, France (Plaziat et al., 1975). 



Material studied. BMNH EE6237. 



Remarks. Differs from H. koninckanus and H. prunella in the 

 relative length of the anterior and posterior petals, and in the angle of 

 divergence of the anterior pair, which is much more acute than in 

 either of those species. 



Family CORASTERIDAE Lambert, in Lambert & Thiery, 1924 



Diagnosis. Paired ambulacra non-petaloid to subpetaloid; more 

 or less flush. Apical system ethmophract. Peristome ovate and 

 downward facing; frontal groove feeble to absent. Peripetalous 

 fasciole only. 



Genus CORASTER Cotteau, 1886 



Diagnosis. Test globose to ovoid with little or no anterior inden- 

 tafion. Apical disc with fourgonopores; test inflated with ambulacral 

 plates relatively tall throughout. Peristome positioned close to the 

 anterior margin and facing forwards. 



Remarks. Distinguished from Homoeaster by the much more 

 anterior position of its mouth, and by its proportionally tall ambulacral 

 plates. The peripetalous fasciole of Coraster also passes much closer 

 to the apical system posteriorly than is the case in Homoeaster. There 

 is considerable variation amongst individuals from the same locality 

 and although many can be placed in one or other of the following 

 species, there is also a good deal of intergradation, making sharp 

 deliniation impossible. 



Coraster vilanovae Cotteau, 1 886 



PI. 11, figs 4-7; Fig. 37a 



1886b Coraster vilanovae Cotteau: 70, pi. 9, figs 1^. 



1975 Coraster vilanovae Cotteau; Plaziat et ai: text-figs 7, 9. 



1995 Coraster vilanovae Cotteau; Gallemi et ai: 269, table 1 . 



Diagnosis. Test subglobular; almost as wide as tall with virtually 

 no frontal sulcus. 



Occurrence. Danian of Erice, Astieso, Larumbe, Navarra. The 

 type locality is the Danian of Les Foies Blanques, Alfas del Pi, 

 Alicante, Spain (Cotteau, 1886b). 



Material studied. BMNH EE6211-17, EE6226-32, MGB 

 37307-18,37325-39,37365. 



PLATE 10 



Figs 1-5 Coraster beneharnicus Seunes. 1888, BMNH EE4823; Thanetian of Larumbe, Navarra. Anterior, lateral, apical, oral and posterior views, xL5. 

 Figs 6-10 Ovulaster reliculatus sp. nov., BMNH EE6074 (holotype), Maastrichtian of Sarasate, Navarra. Apical, oral, lateral, posterior and anterior 



views. x2. 

 Figs 11-13 Coraster beneharnicus Seunes, 1888, BMNH EE6221; Thanetian of Larumbe, Navarra. Lateral, oral and apical views, x2. 



