WEALDEN FORMATIONS. 5 



In the Jurassic Crocodilian (Madrimosaurus, V. Meyer) the coronal ridges of the 

 teeth are more numerous, are smaller at the base of the enamel, and more of the 

 ridges are interrupted than in Goniopholis ; the entire tooth also seems to be shorter and 

 thicker. 



The three vertebrae represented in figs. 1 — 5, PI. II, were obtained by Mr. Holmes 

 from the same bed of Wealden clay, at Cuckfield, as the teeth and scutes, figured in 

 PL III, characteristic of the genus Goniopholis, to which, therefore, I refer them. They 

 correspond with the fourth, fifth, and sixth cervical vertebras of the recent Crocodile, 

 having a parapophysis (p) similar in form, extent, and position, with traces of a short and 

 thick hypapophysis, hy, at the fore part of the under surface; but that surface of the 

 Wealden vertebra is less convex, the whole centrum is relatively broader, and the more 

 important difference of the concavity of the hinder as well as of the fore articular 

 end manifests the distinct family of Crocodilia to which the Goniopholis belongs. The 

 depth of the concavity of these surfaces exceeds that in Teleosaurus. The free surface 

 of the centrum is smooth. The neural arch articulates with the whole length of the 

 centrum by the neurapophysial surfaces of the form shown in fig. 5, np. The neural 

 canal (ib., n) slightly widens behind. Two vertical, venous canals open into the neural one. 

 Fig. 6 is the side-view of a cervical centrum from the Purbeck beds, having the general 

 proportions of those of Goniopholis, but differing in the smaller size of the parapophysis. 

 Figs. 7 — 9 are views of the centrum of a dorsal vertebra of the Purbeck Goniopholis, 

 fig. 9 showing the texture as displayed by a vertical longitudinal section. It is compact 

 or minutely cancellous throughout ; whereas the centrums in the Teleosaurus exhibit a 

 more open, reticulate texture, with a cavity near the centre; this cavity is still larger 

 in Poikilopleuron. 



One of the posterior caudal vertebras of the Cuckfield specimen, after the subsidence 

 of the diapophyses and the great reduction of the zygapophyses, shows the spinous 

 process rising from the hinder part of the neural arch, as at g, PL III. 



The coracoid (ib., h) differs from that of the existing Crocodiles in its greater 

 relative breadth at the neck or part marked h, in the more gradual and minor expansion 

 of its mesial end, and in the more regular convexity of its scapular border. It exhibits 

 the same perforation near this border as in the modern Crocodiles. 



The humerus associated with the remains of Goniopholis from the Wealden of 

 Cuckfield has the usual Crocodilian form and sigmoid flexure. Compared with one from 

 a Crocodilus biporcatus, with the same-sized cervical vertebrae, it is a somewhat thicker 

 and stronger bone. It has a broader and thicker ulnar tuberosity, and the angle 

 at which the process is bent down upon the shaft is less marked, more rounded off. 

 The radial crest is a triangular, compressed ridge, but is not produced beyond four lines 

 from the surface of the shaft ; the distal part of the bone is proportionately thicker 

 antero-posteriorly than in the modern Crocodiles, and the longitudinal, irregular ridges 



