S T A T K OF ILLINOIS DEPARTMENT OF REGISTRATION AND EDUCATION 



BOARD OF NATURAL RESOURCES AND CONSERVATION 

 Wii-LiAM SvivESTER WiiiTE. Chairman ; Thomas Park. Ph.D., Biology; L. L. Sloss. Ph.D., Geology; Roger .Adams, 

 Ph.D., D.Sc. Chftnistry ; Robert H. Anderson, B.S.C.E., Enginetring; Charles E. Olmsted, Ph.D., Forestry 

 W. L. EvERiTT, E.E., Ph.D., Representing the President of the University of Illinois; Delyte W. Morris, Ph.D. 

 President of Southern Illinois I' niversity 



NATURAL HISTORY SURVEY DIVISION, Urbana, Illinois 



SCIENTIFIC AND TECHNICAL STAFF 



Harlow B. Mills, Ph.D., Chief 



Herbert H. Ross, Ph.D., Assistant Chief 



Robert O. Watson, B.S., Assistant to the Chief 



Section of Economic Entomology 



GtOKGE C. Dk KIR, Ph.D.. Principal Scientist and Head 



J. H. Bigger. .\LS., Entomologist 



\V H. Luckmann, Ph.D., Entomologist 



Willis N. Bruce, Ph.D., Entomologist 



JOHN P. Kramer. Ph.D., Associate Entomologist 



Richard J. Dysart, Ph.D., Associate Entomologist 



Ronald H. Meyer, Ph.D., Assistant Entomologist 



Robert D. Palsch, Ph.D., Assistant Entomologist 



Ia.mes E. Appleby. M.S., Assistant Entomologist 



Joseph V . Maddox, M.S., Technical Assistant 



Augustine Okonkwo, M.S., Technical Assistant 



Dannel McCollum, B.A., Technical Assistant 



David Ken.nedy, Technical Assistant 



Sue E. Watkins, Junior Scientific Assistant 



H. B. Petty, Ph.D., Entomologist in Extension* 



Stevenson Moore, III, Ph.D., Associate Entomologist in 



Extension* 

 Clarence E. White, B.S., Technical Assistant in Exten- 

 sion* 

 Amal C. Banerjee, M.S., Research Assistant* 

 Jean G. Wilson, B.A., Research Assistant* 



Section of Applied Botany and Plant Pathology 



J. Cedric Carter, Ph.D., Plant Pathologist and Head 



J. L. Forsberg, Ph.D.. Plant Pathologist 



G. H. Boe«e, ^LS., Associate Plant Pathologist 



Robert A. Evers, Ph.D., Associate Botanist 



Robert Dan Neely, Ph.D., Associate Plant Pathologist 



E. B. HiMELiCK, Ph.D., Associate Plant Pathologist 



Walter Hartstirn, Ph.D., Assistant Plant Pathologist 



D. F. ScHOENEWEiss. Ph.D., Assistant Plant Pathologist 



Peggy Rath, Technical Assistant 



Section of Aquatic Biology 



George W. Bennett, Ph.D., Aquatic Biologist and Head 



William C. Starrett, Ph.D., Aquatic Biologist 



R. W. Larimore, Ph.D., Aquatic Biologist 



David H. Buck, Ph.D., Associate Aquatic Biologist 



Robert C. Hiltibran, Ph.D., Associate Biochemist 



Donald F. Hansen, Ph.D., Associate Aquatic Biologist 



W'.LLiA.M F. Chiiders, M.S., Assistant Aquatic Biologist 



Gary Camenisch, M.A., Technical Assistant 



Ken ^IAcMuRDO, Biochemical Assistant 



Maryfran Martin, Technical Assistant 



Robert D. Cro.mpton, Field Assistant 



Michael J. Duever, Field Assistant 



Charles F. Thoits, III, B.A., Research Associate* 



RoLLiN D. Andrews. Ill, B.S., Field Assistant* 



Larry Bohm, Field Assistant* 



Section of Faunistic Surveys and 

 Insect Identification 



H. II. Russ, Ph.D., Assistant Chief and Head 

 Milton W. Sanderson, Ph.D., Taxonomist 

 Lewis J. Stannard, Jr., Ph.D., Taxonomist 

 Philip W. Smith. Ph.D., Taxonomist 

 Leo.nora K. Gloyd, M.S., Assistant Taxonomist 

 H. B. Cunningham, Ph.D., Assistant Taxonomist 

 Bern ice Sweeney, Technical Assistant 

 George L. Rotramel, Technical Assistant 

 Marvin E. Braasch, Laboratory Assistant 

 John D. Unzicker, B.S., Research Assistant* 

 TosHio Yamamoto, B.S., Research Assistant* 



Section of Wildlife Research 



Glen C. Sanderson, Ph.D., Wildlife Specialist and Hea> 



F. C. Bellrose, B.S., Wildlife Specialist 



H. C. Hanson, Ph.D., Wildlife Specialist 



Richard R. Graber, Ph.D.. Wildlife Specialist 



Ronald F. Labisky, ALS., Associate Wildlife Specialist 



William R. Edwards, M.S., Associate Wildlife Specialii 



William W. Cochran, Jr., Assistant Wildlife Speciali: 



Nancy Kott. B..^., Technical Assistant 



Richard Bartholomew, B.S., Technical Assistant 



Howard Crum, Jr., Field Assistant 



John E. Warnock, Ph.D., Research Associate* 



Jack A. Ellis, M.S.. Research Associate* 



Bobbie Jo Verts, M.S., Research Associate* 



Ralph J. Ellis, \I.S., Research Associate* ' 



Gerald G. Montgomery. M.S., Research Associate* 



William L. Anderson. B.S., Research Associate* 



George B. Joselyn, \LS., Research Assistant* 



Steven L. Wunderle, B.S., Research Assistant* 



Richard D. .'\ndrews, M.S., Field Mammalogist* • 



Gerald L. Storm, M.S., Field Ecologist* 



Terence R. Troughton, B.S., Field Biologist* 



David A. Casteel, B.S., Project Assistant* 



Ronald L. Westemeier, B.S., Project Assistant* 



Keith P. Dauphin, Project Assistant* 



Section of Publications and Public Relations 



James S. Avars, B.S., Technical Editor and Head 

 Blanche P. Young, B.A., Associate Technical Editor 

 Marguerite AI. Verley, M.A., Assistant Technical Edit 

 WiLMER D. Zehr, Assistant Technical Photographer 



Technical Library 



Doris F. Dodds, B.A., M.S.L.S., Technical Librarian 

 Patricia F. Stenstrom, B.A., M.S.L.S., Assistant Tec 

 nical Librarian 



CONSULTANTS: Herpetolocy, Hobart M. Smith, Ph.D., Professor of Zoology, University of Illinois ; Pakasitoi-oc 

 Norman D. Levine, Ph.D., Professor of Veterinary Parasitology and of Veterinary Research, University of Illino\ 

 W'lLDLiFE Research, Willard D. Klimstra, Ph.D.. Professor of Zoology and Director of Co-operative Wildlife / 

 search, Southern Illinois University; Statistics, Horace W. Norton, Ph.D., Professor of Agricultural Statisti't 

 Design and Analysis, University of Illinois. 



♦Employed on co-operative projects with one of several agencies: University of Illinois, Illinois Agricultural Ext( 

 sion Service, Illinois Department of Conservation, National Science Foundation, United States Department of A] 

 culture. United States Fish and Wildlife Service, United States Public Health Service, and others. 



This paper is printed by authority of the State of Illinois, IRS Ch. 127, Par. 58:12. It is a contribution from 

 Section of Applied Botany and Plant Pathology of the Illinois Natural History Survey. 



(88590— lOM— 2-64) c^^^z 



