1847.] OWEN ON ENGLISH FOSSIL REMAINS OF MEGACEROS. 45 
are occupied by a slender but long accessory column of enamelled 
dentine (p); in Megaceros this column is represented in those teeth 
by a pyramidal tubercle at the bottom of the interspace (fig. 2. p). 
In molars, therefore, which have been worn down to the same ex- 
tent as in those of the fossil Bovine and the Megaceros represented in 
Fig. 3. Bos primigenius. 
Fig. 4. Megaceros. 
figs. 3 & 4, the column, p, increases the complexity of the grmdmg 
surface in the one, but not in the other. The third lobe, o!, in the last 
molar, M 3, of Bovines is terminated by a similar enamelled column or 
‘columella,’ g, fig. 1, which is not present in the Megaceros. The 
minor differences of the grinding surface are sufficiently illustrated by 
the figures, which may aid future collectors in determining similar 
evidences of additional examples of Megaceros in British pleistocene 
strata. 
The teeth of the Bos longifrons are readily distmguished by their 
smaller size. This species of Bos co-existed with Megaceros in 
Treland, and in England with Megaceros, Rhinoceros, Elephas, 
Hyena, &c. 
Remains of Bos longifrons have, however, also been found in the bog 
itself, as well as in the subjacent marl, in Ireland ; but I have not, 
as yet, had any trustworthy and authentic evidence of the discovery 
of undisturbed remains of Megaceros in the peat-bog above the shell- 
marl in any part of Ireland. 
Remains of Bos longifrons have been found in ancient places of 
sepulture, and so associated with British and Roman remains, as to 
leave little doubt of its having survived, as a species, many of the 
mammals with which it was associated during the pleistocene period 
in geology. Whether the Megaceros, likewise, continued to- exist 
until the ‘Human period,’ is a question on which I have never enter- 
tained or expressed a decided opinion, knowing the uncertainty of 
negative evidence. But, as yet, I am bound to state that no good 
evidence to the contrary, z.e. no evidence demonstrating the co-ex- 
