HATCHER : OLTGOCENE CAN ID ^ 67 



cene formation at Sansans, France. The reference of this material to the genus 

 Amphicyon seems to have been based entirely upon the dental formula and is now 

 known to be erroneous, as has been shown by Scott. The chief [generic distinc- 

 tions between Daphcemis Leidy and Amphicyon Lartet in so far as they are at 

 present known are to be found in the relative size of the premolars, structure 

 of canines, and position of the superior third tubercular molar. The canines 

 of Daphoenus are without either anterior or posterior cutting edges, while these 

 are present in Amphicyon. The premolars are reduced in size in Amphicyon, 

 while those of Daphoenus show little or no reduction. In Daphoenus the superior 

 third tubercular molar is pushed inward and aligned with the internal cones 

 of the preceding molars, while in Amphicyon this tooth occupies a more external 

 position. 



In 1869,^ as Amphicyon vetus, Leidy gave a much more complete description of 

 the material which had formed the type of his previously proposed genus Daph- 

 oenus. Among the more important additional characters then enumerated by him 

 the following may be mentioned as being perhaps of generic importance. 



1. Presence of long, strong and high sagittal crest. 



2. Auditory buUse small. 



3. M.^ small, transversely oval, with two tubercles and one root. 



In 1898^ Scott confirmed most of the observations made by Leidy and added a 

 great number of new characters relating not only to the skull and dentition, but to 

 other portions of the skeleton as well. In this paper Professor Scott gave the first 

 adequate account of the osteology of the genus Daphoenus and pointed out its dis- 

 tinction from Amphicyon. On account of the incomplete nature of the material at 

 his command Scott's description was necessarily based on the remains of several 

 more or less fragmentary skeletons belonging to different species. Moreover, as we 

 shall show later, it is not at all certain that some of the material described by Scott 

 does not pertain to a distinct genus. 



In 1899 Wortman and Matthew" characterized the genus Daphcemis Leidy as 

 follows : " Upper molars transversely unsymmetrical (paracone more external than 

 metacone). M- oval, aligned with inner cusps of anterior molars. Heels of lower 

 molars low-ridged, with low entoconid crest, nig a convex nub." 



Combining the characters above noted with those shown by the material under 

 discussion, the genus Daphoenus may be distinguished by the following cranial and 

 dental characters. 



*See Journ. Acad. Sci. of Phila., 1869, pp. 31-36. 

 sSee Trans. Am. Phil. Soc, Vol. XIX., pp. 325-415. 

 6 See Bull. Am. Mus. Nat. Hist., Vol. XII., p. 129. 



