136 DK. A. SMITH WOODWAKD ON VERTEBEATE [May I907, 



the ginglymoid end of the quadrate bone. In shape and relation- 

 ship it agrees precisely with the articular bone of Macropoma 

 (PI. VIII, fig. 7, art.). Behind the articular there is at least one 

 other stout ossification of the meckelian cartilage, which extends 

 backward even beyond the angular and is visible from the outer 

 face of the mandible (PI. VIII, fig. 2, m.), where its thickened 

 upper portion is conspicuous and peculiar, leaving a pit between 

 this thickening and the posterier edge of the angular element (ag.). 

 The same arrangement is also seen in Macropoma (PI. VIII, fig. 8). 

 Within the articular and the hinder ossification just mentioned, the 

 inner face of the mandibular ramus is formed by one or more thick 

 splenial bones, of which the precise extent forward is unknown. 

 A much elevaced coronoid process is formed by a separate bone, 

 articulating with the outer face of the splenial in a narrow facette 

 which is deepest behind (PI. VIII, fig. 3, /). This element, which 

 corresponds with the ascending process of the splenial in Polypterus, 

 is seen from the outer aspect on the right side of the type-skull 

 (PI. VII, fig. 1, co.), and is detached on the left side of the same 

 specimen, thus displaying also its inner aspect (PI. VIII, fig. 3). 

 It is triangular in shape, with the middle of its antero-superior 

 edge thickened, indented, and projecting outward ; while its sig- 

 moidally-bent inner face is covered with minute tubercular teeth 

 like those of the pterygoid bones. The anterior part of the mandible 

 remains to be discovered, but it probably bore small conical sculp- 

 tured teeth rather longer than those of the pterygoid plates, 

 because a few such teeth are scattered on the inner side of the left 

 mandibular ramus in the type-specimen. 



Broken remains of the characteristic branchial arches project 

 behind the cranium in the type-specimen (PI. VII, fig. 1, 6r.), 

 and one detached ceratobranchial, from Mapelle Quarry, is especially 

 well preserved. This bone measures 48 centimetres in length, 

 tapers at each end, is laterally much compressed, and exhibits the 

 usual deep groove posteriorly. 



One of the gular plates found with the type-skull is nearly com- 

 plete (PI. VIII, fig. 5), and its length equals about 3J times 

 its maximum width. Its outer face is raised into a smooth longi- 

 tudinal ridge, slightly curved, near the outer border of the bone ; 

 and from this radiates a fine ridged ornament, which is in places 

 feeble, in places reticular. 



Isolated portions of the operculum, undoubtedly of the same 

 species, indicate that this bone was shaped as usual in Coelacanths, 

 but much produced downward at its antero-inferior angle. It is 

 completely covered with radiating ridges, which resemble those of 

 the head-bones in being destitute of enamel and nearly smooth ; 

 and they radiate from the point of suspension, where they are 

 fewest and tend most towards reticulation (PI. VIII, fig. 6). 



An isolated clavicle from Itacaranha (B. M., No. P 10061), large 

 enough to have belonged to the type-specimen, is typically Ccela- 

 canth in its slightly-expanded, rounded upper end, and in the small 



