H.. J. Clark on Lucernaria. 351 
the muscular layer and the inner wall, which forms the imme- 
diate parietes of the digestive cavity, a thick layer of gelatini- 
form substance intervenes, and its presence naturally suggests 
the inquiry, how are the eggs or sperm to escape into the diges- 
tive cavity, as they are known to do? The spermatic particles 
Thave observed frequently escaping directly through the outer 
wall into the ocean, and I have seen them, with the broadest end 
Out, projecting like bundles of hairs from the cavity of the matrix 
through the apertures in the outer wall. When the reprodue- 
ive material is fully ripe, the inner wall, with the gelatiniform 
layer, and the museular layer as far as it includes the material 
in question, splits off from the outer wall along two lines corres- 
pacing to the two borders of the generative organ, and hangs 
oosely, in ribbons, in the digestive cavity. From the newly- 
formed raw face of these ribbons the eggs or spermatic particles 
€scape into the main chamber of the disc. This I take to be 
the universal rule, and such the type of genitalia among all 
Steganophthalmata; a structure totally unlike that of Lucerna- 
Ma, mm which the nner wall alone is concerned in the highly 
complicated reproductive organs. 
+ 4ssing now to the consideration of the muscular system, I 
will call your attention to the four white, slender columns which 
alternate with the four dark tubes which are imbedded in the 
Selatiniform substance of the peduncle. Sars was the first to 
indicate the true nature of these columns, and he rightly called 
them muscular cords. They extend from the base of the pedun- 
cle to the base of the proboscis, coursing along just beneath the 
Suter wall, but still within the gelatiniform substance, until they 
Teach the upper third of the peduncle, and then gradually ap- 
Proximating the axial line, they meet the inner wall of the disc 
Just below the base of the proboscis, and thence they pass along 
still beneath this wall, for a short distance, and, finally each one 
fnters the oral side of the disc at the inner or axial end of the 
_ partition, At this point, each muscular column expands and 
ue a fan-shaped layer just beneath the outer wall, and extends 
laterally so as to occupy the whole space between the two halves 
of a genital. At the distal end, this layer diverges right and 
left of the partition into a broad muscular band which borders 
the dise, and, eventually, is distributed in ridges or cords beneath 
the outer wall of the tentacles and the auricles, At the inner 
_ &nd of the partition, the muscular layer also passes into the base 
_ Of the proboscis, and forms a stratum itnmnieavataly beneath the 
ater wall. At four equidistant points, alternating with the 
Partitions and genitals, and opposite the four corners of the 
_ proboscis, there is a weaker muscular layer, which occupies the 
_ Same relative position in regard to the outer walls as does the 
_ *Wonger system of muscles first mentioned. On the one hand, 
