158 



On the Reproductive Functional 



[No. 



the results of fertilisation with its own pollen. Fourthly, in accordance 

 with recognised systematic affinities, we find the following descending 

 scale of sterilisation resulting from the unions of V. nigrum, V. virgatum 

 and V. thapsiforme with the V. lychnitis. Thus with the pollen of 

 V. nigrum, the average fertility of V. lychnitis, lutea, is 80, with 

 that of V. virgatum 58, and with that of V. thapsiforme 46, rela- 

 tively, in each instance, to 100, the product of fertilisation by its 

 own pollen. A similar accordance is observable between the functional 

 and systematical relations of V. hlattaria and V. thapsus with the 

 V. lychnitis. In the unions, however, of V. phoeniceum and varieties 

 with the V. lychnitis, no such accordance is observable. The different 

 unions vary greatly in the degree of fertility inter se, and judging 

 indeed by the relative functional potency of the pollen of the three 

 varieties on the stigmas of V, lychnitis, the different results are 

 comparable with those from distinct species, and would cause their 

 interpolation into systematically considered false positions, showing 

 us that the functional and systematic affinities of the species of a 

 genus are by no means strictly co-ordinated. 



m «■ 



.A 



p 





03 



By calcu- 









& 





CD 

 CD 



lation. 







«M 



co 





CO 





The 

 comparative 



Table 5. — Pure and Mixed Unions 



CO 



<3 







CO 



d 





CO 



of Verbascum hlattaria, L. var. 



g 



CO 



H3 

 CD 



Pi 





CD 



fertility of 



alba of gardens. 



=0 . 





CD 

 to 



6C ° 



^-( CD 



CD 

 00 



the different 





o <o 



o§ 



o 



03 jL, 



CD £ 



°1 



O 



unions. 





d " 



6 ^ 



6 



l> —' 



ft cT 



d 







ft 



ft 



ft 



•4 



o 



ft 





No. 







1. Verbascum hlattaria, L. var. 



















alba of gardens by own 



















pollen, 



8 



8 



438 



55 



20 



1095 



1000 





2. V. hlattaria, alba by pollen 



















of V. hlattaria, lutea,h., . 



6 



5 



217 



43 



20 



868 





792.6 



3. V. hlattaria, alba by pollen 



















of V. thapsus, lutea, L., 



6 



2 



36 



18 



20 



360 





328.7 



4. V. hlattaria, alba by pollen 



















of V. thapsus, L. var. alba 



















of gardens, 



6 



4 



95 



24 



20 



475 





433.7 



5. V. hlattaria, alha by pollen 



















of V. lychnitis, lutea L., . 



8 



5 



65 



13 



20 



260 





237.4 



6. V. hlattaria, alba by pollen 



















of V. lychnitis, L. var. alba 



















of gardens, 



6 



4 



79 



20 



20 



395 





360.7 



