SCLEROTIA OF UNKNOWN AFFINITY. 267 



neck of living Hhinanthus minor; these bodies begin their 



development in the cambium and bark, v?hich they kill ; after- 

 wards the wood itself mav be attacked. 



Sclerotinia with Botrytis-conidia/ 



Scl. Fuckeliana De Bary. This Sclerotinia is distinguished 

 from all preceding ones by its passing through a Botrytis- 

 conidia stage {Botrytis cinerea). If conidia are sown out on 

 plum-juice gelatine, there appear within fourteen to twenty-one 

 days round groups of sclerotia, which soon 

 give rise to conidia. From such artificially- 

 reared sclerotia I have never succeeded in 

 getting the Peziza-tmit, so easily cultivated 

 from sclerotia gathered in the open-air (e.g. 

 from vine leaves).^ Thus the actual proof 

 that Scl. Fuckeliamt and Botrytis cinerea are Fjlk°iimm.'~m^-mt'^ 

 stages in the life of the same fungus is not soieroUa^' (v.^^iubeuf 

 reached by this experiment.^ The two forms ^'"'^ 

 are, however, very frequently met together. 



The sclerotia of Scl. Fiickeliana are produced in the mesophyll 

 of the leaves, also in the parenchyma and epidermis of the 

 host-plants, but never in the wood. Peziza-hmts. with flat 

 apothecia are produced from them. Sclerotia are found in vine 

 leaves and over-ripe grapes (Fig. 140), especially of the Eiesling, 

 Orleans, and Sylvaner varieties.* Other plants and fruits may 

 also be attacked. Diseased parts become brown from the 

 effects of the parasitic mycelium, and die off. The mycelium 

 can only live parasitic after it has been strengthened by a pre- 

 vious saprophytic existence. Ascospores are thus unable to 

 effect direct infection. The ^o^/'^/^is-conidia seem, however, 

 capable of directly infecting a host-plant, at least I have always 

 succeeded in infecting Conifers successfully with the conidial 

 form Botrytis Douglasii. 



'See also Botrytis amongst the "Fungi imperfecti." 



^ Brefeld, Heft I v. , p. 129, and x. , p. 315 ; Tubeuf , Beitrage z. Kenntniss d. 

 Baumkrankheiten, 1888. 



^Zopf. (Die Pilze, p. 742) states that Peztza-fruits may be reared from these 

 sclerotia after they have rested a year. 



^ MuUer-Thurgau, "Die Edelfaule d. Trauben." Landwirth. Jahrbuch, 1888 

 (Ref. in Botan. Centralblatt, xxxv., 1888, p. 94). 



