EXTERNAL PABTS OF BIRDS. — THE FEET. 



129 



aud functional efficiency. In most Passeres it is virtually provided with a special muscle f(jr 

 independent movement, so that it may be perfectly apposahle to the other toes collectively, 

 just as our thumb may be brought against the tip of any finger. In general, it shortens as it 

 rises on the metatarsus; and probably in no bird in which it is truly elevated is it as long as 

 the shortest anterior toe. It is short, barely touching the ground, iu most wading birds; 

 shorter still in some swimmers, as the gulls, v\-here it is probably fuiictionless ; it is incom- 

 plete in one genus of gulls (Rissa), where it bears no perfect claw ; it has only one phalanx 

 and is represented only by a short immovable claw in the petrels (Procellariidije) ; it disappears 

 in the birds named in the last paragraph but two above, and in some cjtliers. It is never actu- 

 ally soldered with any other toe, for any noticeable distance ; but it is webbed to the base of the 

 inner toe in the loons (Colijmhus), and to the whole length of the toe in all the Steganopodes 

 (fig. 52). It may also be independently webbed; that 'm, be provided with a separate flap or 

 lobe of free membrane. This lobation of the hallux is seen iu all our sea-ducks and mergansers 

 {Fuligulina: and Meryina), and iu all the truly lobe-footed birds, as coots (Fulica), grebes 

 (Podicipedidce) and phalaropes {Phalaropodida) . The modes of union of the anterior toes 

 with one another may be finally considered under the head of the 



Three leading Modiflcations of the Avian Foot. — Birds' feet are modelled, on the 

 whole, upou one or another of three plans, furuisliiug as mscay types of structure ; which 

 types, though they run into one another, and each is variously modified, may readily be appre- 

 ciated. These plans are the perching or insessoriul, the walking or wading, cursorial or 

 grallatorkd, and the swimming or natatorial — iu fact, so well distinguished are they, that 

 carinate birds have even been primarily divided iuto groups corresponding to these three 

 evidences of physioh igical adaptation of the structure of the Avian ^jes. Independently of the 

 number and positicjn of the digits, the plans are pretty well indicated by the method of union 

 of the toes, or their entire lack of union. 1. The insessorial type, (n) In order to make a foot 

 the most of a hand, that is, to fit it best for that grasping function which the perching of 

 birds upon trees and bushes requires, it is requisite that the digits should be as free and 

 movable as possible, and that the hind one should be perfectly apposable to the others. 

 Compare the human hand, for example, with the foot, and observe the [lerfection secured by 

 the perfect freedom of the fingers and especially the appositeness of the thumb. In the most 

 accomplished insessorial foot, the front toes are cleft to the base, or only coherent to a very 

 slight extent ; the hind toe is completely incumbent, and as long and flexible as the rest. Our 

 thrushes (Tardidce) probably show as complete cleavage 

 as is ever seen, practically as much as that of the 

 human fingers ; the cleft between the inner and middle 

 toe being to the very base, while the outer is only joined 

 to the middle for about the length of its own basal 

 joint. This is the typical passerine foot (figs. H6, 37, 

 42, iii). There may be somewhat more cohesion of 

 the toes at base, as in the wrens, titmice, creepers, 

 vireos, etc., without, however, obscuring the true pas- 

 serine character. As regards this matter, the point is, 

 that when the toes are united at all, it is by their actual 

 cohesion there, not by movable webbing. Besides the 

 typical passerine, there are several other modifications 

 of the insessorial foot. (6) Thus a kingfisher shows 

 what is called a syndactyle or syngnesious (Gr. crvv, .mn, together; yvrja-ws, gnesios, relating to 

 way of birth) fijot (fig. 44), where the outer and middle toes cohere for most of their extent and 

 have a broad sole in common. It is a degradation of the insessorial foot, and not a common 



Pigs. 42, 43. — Typical passerine feet. 

 (The right-hand fig. is Ptecirophants lappo- 

 nicns, nat. size.) 



