170 BULLETIN 50, UNITED STATES NATIONAL MUSEUM 



Auk, XXV, 1908, 432 (Bellinghara Bay region. Wash. ; common except on higher 

 mountains). — Dawson and Bowles, Birds Washington, ii, 1909, 587, part 

 (habits; distr.).— (?) Willett, Condor, xvi, 1914, 89 (doubtful record for 

 Sitka, Alaska; apparently=Westminster, British Columbia).— Bum-eigh, Auk, 

 xlvi, 1929, 510 (Tacoma, Wash.; breeding habits).— American Ornithologists' 

 Union, Check-list, ed. 4, 1931, 82, part.— Miller, Lumley, and Hall, Mur- 

 relet, xvi, 1935, 57 (San Juan Islands, Wash.).— Kitchin, Murrelet, xx, 1939, 

 30 (Mount Ranier National Park). — Hellmayr and Conqver, Cat. Birds Amer., 

 i. No. 1, 1942, 218 part. 



Bonasa sabinei Elliot, Monogr. Tetraonidae, 1865, pi. 3 and text, part. 



Bonasa umbellus togata Edson, Auk, xxv, 1908, 432 (Bellingham Bay region. Wash.; 

 in the mountains). 



Bonasa umbellus castaneus Aldrich and Friedmann, Condor, xlv, 1943, 95 (Soleduck 

 River, OljTTipic Mountains, Washington; tax.; crit. ; descr. ; distr.). 



BONASA UMBELLUS BRUNNESCENS Conover 



Vancouver Island Ruffed Grouse 



Adult (brown phase). — Similar to that of Bonasa umbellus umbellus 

 but darker, more brownish, less rufescent (darker, more brownish than 

 sabini also) ; general color of the upperparts between Prout's brown and 

 Dresden brown, tail dull ochraceous-umber ; underparts heavily barred 

 with grayish ochraceous-umber and washed extensively with tawny-buff. 



Adult (gray phase). — Similar to the brown phase, but the top of head, 

 neck, back, rump, and upper tail coverts mixed and vermiculated with 

 dark smoke gray ; rectrices dark smoke gray barred and vermiculated with 

 black and without any brownish tinge (the heavy black wavy bars are 

 single in this race in both phases, while in sabini and castanea they 

 are double with a pale ochraceous band in between them) ; underparts 

 as in the brown phase but much less washed with buflEy. 



Juvenal.— Svcmlar to that of B. u. umbellus but darker brown (darker 

 than Juvenal monticola also) ; above, cinnamon-brown to Prout's brown 

 (as opposed to sayal brown in the nominate form) ; below, the ventral 

 barrings darker — dusky Dresden brown. 



Downy young. — None seen. 



Adult male.— Wing 179-189 (183.7) ; tail 144-157 (148.6) ; culmen 

 from base 26.6-28.3 (27.6) ; tarsus 44.0-46.8 (45.6) ; middle toe without 

 claw 40-41 (40.3) ; unfeathered part of tarsus 24.7-29 (26.7 mm.)." 



Adult female.— Wing 173-181 (176.3) ; tail 124-134 (128.4) ; culmen 

 from base 24.4-27.4 (26) ; tarsus 41.8-45.2 (43.0) ; middle toe without 

 claw 37-39.9 (38.4) ; unfeathered part of tarsus 20.5-28 (25.3 mm.).^« 



Range. — Subclimax woodland of the cedar-hemlock association in the 

 moist coniferous forest biome (Transition Life Zone) ; on Vancouver 

 Island, British Columbia, and the adjoining mainland from the vicinity 



" Six specimens including the type. 

 " Six specimens. 



