18 EDWARD T. BROWNE. 
The jelly of the umbrella is not only very thick, but very firm. There is no 
apical depression in the wall of the umbrella, as found in S. nawarchus. The margin 
of the umbrella is divided by vrooves, which are opposite the tentacles, into 
small lobes. 
The mouth has four short perradial lips, and the margin is arranged in slight 
folds. Just inside the mouth there are four interradial, endodermal processes 
somewhat triangular in shape. On the closing of the mouth these processes meet 
and close the entrance to the stomach. One specimen is, however, abnormal, and its 
mouth has seven pointed lips. It has five longitudinal ridges on the stomach. 
The most interesting features in this Medusa are the gonads and their position 
with regard to the stomach. The stomachs of two specimens were cut into transverse 
sections. One series (Plate II., fig. 4) shows ripe spermaries, and the spermatozoa 
in the process of being discharged; the other series (Plate II., fig. 5) shows the 
condition of the stomach after the gonads have been discharged. 
On the outside of the stomach there are four perradial longitudinal bands (fig. 2) 
which slightly project as ridges. Inside each ridge runs a canal-like cavity lined with 
endoderm (fig. 4). These canals branch out from the interior of the “ mesenteries,” 
and are in direct communication with the top of the stomach and also with the radial 
canals. They run down the wall of the stomach nearly to the mouth, and there 
terminate blindly, without any communication with the exterior. 
Between the perradial bands on the outside of the stomach there are numerous 
small holes (fig. 2). The shape of the holes varies in the different specimens, and they 
may be either circular, oval, or somewhat quadrangular. The holes are arranged in a 
single row on both sides of the perradial bands, and a few occupy the interradial spaces 
in the upper part of the stomach. Within these holes, or protruding from them, is a 
whitish flocculent substance, which is composed of spermatozoa. 
The sections show very clearly that the growth of the gonads has converted the 
stomach into a reproductive organ, and that its function as a stomach has ceased. 
There is practically no cavity for the reception and digestion of food, for although 
a very small cavity does exist in the centre of the stomach (fig. 4), it is not in 
communication with the mouth. In the lower part of the stomach the endoderm forms 
a solid mass in the centre. 
The spermaries form globular or spherical masses encased in a very thin 
membrane which lies next to the endoderm of the stomach. The endoderm, stained 
with hematoxylin, in the sections, has the appearance of a mosaic pavement in 
different tints of blue. The cells have not a well-defined wall and are filled with a 
rather dense homogeneous cytoplasm. The preservation is not good for cytological 
details, and it must be borne in mind that the specimens were merely preserved for the 
determination of the species. 
It is unfortunate that there are no intermediate stages of this Medusa in the 
collection for the elucidation of the development of the gonads. The four perradial 
