MEDUSA. 53 
the tentacular series, regularly alternating with the former. The tentacular canals are 
evidently not of the same age as the rhopaliar canals, but of a slightly later growth. 
The tentacles are just beginning to make their appearance, and are indicated 
either by bulb-like buds, by tapering elongated buds, or by minute tentacles. Four 
tentacular buds varying in size and age are found in the smallest specimens, and as 
the Medusa grows twelve more buds develop, making the total up to sixteen. It is 
clear that the tentacles do not all develop at the same time, but at irregular intervals 
and apparently in no definite order. Ultimately the full number is reached, and 
corresponds to the number of sense organs. 
It is quite probable that there is a still younger ephyra stage, which is not 
represented in the collection—a stage with only sixteen rhopaliar canals, and without 
the tentacular canals. 
The stomach is very small, circular in outline, and four gastric filaments are 
visible inside. The filaments increase in number as the Medusa grows. In the early 
stages one filament in each group is much longer than the others, and this is 
probably the primary one. The mouth is simply a large opening, without any definite 
lips or arms, which appear later. 
The ex-umbrella is covered with small clusters of nematocysts. In the later 
stages the nematocysts are confined to the aboral side of the marginal lobes. 
The circular canal is formed by outgrowths from the,radial canals (Plate VL, 
fig. 2), and is evidently formed just before the branches of the rhopaliar canals begin 
to develop. . 
The Meta-ephyra stage.—The normal appearance of this stage has been very 
well figured by Maas (1908, Plate II., fig. 2). It may be distinguished by all the 
rhopaliar canals possessing two opposite lateral branches, which lead into the circular 
canal. The tentacular canals remain unbranched, as in the earlier stages. The 
branching of the rhopaliar canals in the ‘ Discovery’ specimens is rather irregular, and 
there is a want of uniformity in the pattern. The number of tentacles or tentacular 
buds present is very variable and is not correlated with the size of the umbrella, which 
at this stage is 15 to 25 mm. in diameter. 
Variation.—Among the ephyra and meta-ephyra stages fourteen specimens are 
sufficiently perfect for counting the number of rhopalia or rhopaliar canals. One 
specimen has eleven rhopalia, four have fourteen rhopalia, two have fifteen rhopalia, 
and seven have the normal number of sixteen. 
The early stages were mostly taken by the ‘ Discovery’ during April and May. 
As meta-ephyra stages were taken at the end of March and ephyra stages in May and 
June, there is no clue given as to the breeding season of this Medusa. The ephyra and 
meta-ephyra stages were found by the ‘Gauss’ in January and March. Vanhéffen’s 
account of these stages is based upon nine specimens, 4 to 22 mm. in diameter. 
The Adult.—The adult specimens were all placed in one jar and labelled “‘ Winter 
Quarters, various dates, 1903. Chromic-formalin.” They were afterwards transferred 
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