THE PHANEROGAMS OF THE JUAN FERNANDEZ ISLANDS 1 69 



Euphrasia L. 



*io3. E. formosissima nov. spec. — Plate 15, fig. 1, Plate 20, fig. 6—j; 

 text fig. 23. 



Eueuphrasia pluriennis lignosa, plerumque 15 — 20, interdum usque ad 70 cm 

 alta, ad radices Pernettyae rigidae parasitica. Truucus primarius cylindrico- 

 tetragonus, ad 6 mm crassus, cortice obscure fusco inferne glabrato, fere a basi 

 opposite ramosus, ramis ramulosis ramulis iterum ramosis, junioribus distinctius 

 tetragonis bifariam hirsutis, pilis crassis plerumque bicellularibus. Folia carno- 

 sula late — anguste ovata, 8 — 18 (rarius ad 20 et ultra) mm longa et 5 — 8 mm lata, 

 basi in petiolum 2 — 3 mm longum subiter vel sensim angustata, margine leviter 

 revoluta dentibus humilibus latissime truncatis utrinque 3—5 (plerumque 4) 

 instructa, supra subglabra, parcissime glandulosa sed secus nervos pilosiuscula, 

 subtus in parte inferiore ad nervos parce pilosa vel subglabra, infra marginem 

 et inter nervos incrassatos reticulo glandularum humilium capite ferrugineo 

 pulchre notata. Bracteae quam folia caulina minores ac latiores. Flores ad 

 apicem caulis ramorumque breviter sed densiuscule spicati, sessiles. Calyx pro 

 genere sat typicus, lobis lineari-subspathulatis apice incrassatis, extus nervis 

 exceptis subglaber, intus secus nervos pilosus pilis unicellularibus et glandulis 

 ferrugineis commixtis, 5 — 6 mm longus, profunde partitus. Corolla alba 10 — 13 

 mm longa, tubo 4 — 5 mm longo ad finem anthesis haud elongato, subglabro, 

 intus inferne obscure luteo; labium superius extus pilosum, 6 — 7 mm longum 

 et 5 — 6 latum, bifidum lobulis reflexis; labium inferius glabrum 7 — 8 mm longum 

 et ad 10 — 11 mm latum, in centro macula magna flavo-aurantiaca distincte cir- 

 cumscripta notatum. Filamenta staminum posticorum sub 3 mm, anticorum circ. 

 3,5 mm longa, inferne lutea, dein viridescentia, antherae ad 1,4 mm longae, 

 ferrugineae, subglabrae-glabrae, duorum staminum posticorum loculus unus paulo 

 longius calcaratus. Stylus ad 9 mm longus, dilute viridis, parce pilosus, exsertus; 

 stigma alboviride capitatum. Capsula matura valvis obovatis minute emarginatis 

 et abrupte apiculatis, superne setulosa, 2,8—3 mm longa et 2 — 2,1 mm lata. 

 Semina oblique fusiformia I — 1,1 mm longa straminea, testa longitudinaliter 

 sulcata et inter juga transverse sculpta. 



Masafuera: in the higher parts, not uncommon in the alpine region. 

 Ridges above Q. Sanchez (fl. 25 /2 17, no. 386 b); north of the Casas gorge from 

 800 to 1400 m (fl.-fr. u /2— 7 /3 17, no. 386); C. Atravesado, common; C. del 

 Barril, from 750 m (fl. Va 17, no. 547). 



A rather interesting discovery. Two types of Euphrasia are known from 

 the southern hemisphere, sect. Trifidae from Chile and sect. Etieuphrasia sub- 

 sect. Australes from Australia and New Zealand. The new species has nothing 

 to do with the South American species, and as far as I can see, it is not 

 nearly related to the Australes, but belongs to the Semicalcaratae, known before 

 only from the northern hemisphere. Within this section E. formosissima is the 

 only perennial species. Among boreal ones it might be compared with E, 

 grandiflora Hochst. (Azores), or with species from Eastern Asia. It lies near 

 at hand to regard it as one of the older endemics in the islands, as it differs 



