ALIMENTARY SYSTEM. 407 
non-glandular, and those of the second type are often somewhat 
eye-like, with pigment layer, reflector, lens, and diaphragm, or with 
some of these structures. . 
Alimentary system.—The cuttlefish eats food which 
requires tearing and chewing, and this is effected by the 
chitinous jaws worked by strong muscles, and by the 
toothed radula moving on a muscular cushion. The mouth 
lies in the midst of the arms, bordered by a circular lip, and 
opens into a large pharynx or 
buccal cavity (cf. the snail). The p-Z 
narrow gullet passes through the Ch 
ganglionic mass, and leads into st, 
the globular stomach, lying near 4 
the dorsal end of the body. The 
stomach is followed by a cecum 
or pyloric sac, and the intestine 
curves headwards again, to end 
far forward in the mantle cavity. 
There do not seem to be any 
glands on the walls of the food g 
canal; the stomach has a hard = § 
cuticle ; the digestion which takes 
place there must therefore be 
due to the digestive juices of the 
glandular annexes. Of these the 
Fic. 218.—Diagram of the 
structure of Sepza.— Mainly 
most important is usually called 
the liver; it is bilobed, and lies 
in front of the stomach, attached 
to the cesophagus. Its two ducts 
conduct the digestive juice to 
the region where the stomach, 
“pyloric sac,” and intestine 
meet; and these ducts are 
fringed by numerous vascular 
and glandular appendages, which 
are called “ pancreatic,” 
after Pelseneer. 
a., Eight short arms around mouth ; 
Z.a., one of the two long arms; 
b., beak of the mouth; c.g., cere- 
bral ganglia, with commissures 
to the others; £., eye; 2, 
gullet; ag., digestive gland (the 
“salivary glands” are not repre- 
sented); sz., stomach; @., anus} 
she, shell-sac with sepiostaire } 
&, kidney; &., reproductive 
organ; 4r.., branchial heart ; 
fy a zill; 2.6., ink-bag; 712.¢., 
mantle cavity; “, funnel. 
and arise from the wall of the 
unpaired portion of the nephridia. 
Far forward, in front 
of the large digestive gland, lie two small white glands 
on each side of the gullet, with ducts which open into the 
mouth (cf. the “salivary glands” of the snail). A diastatic 
ferment has been proved in the salivary secretion of 
