APPENDIX I 



As for this unfinished work, suggestive outlines left for 

 others to fill in, Professor Howes writes to me in October 

 1899:— 



Concerning the papers at S.K. which, as part of the contents 

 of your father's book-shelves, were given by him to the College, 

 and now are arranged, numbered, and registered in order for 

 use, there is evidence that in 1858 he, with his needles and eye- 

 glass, had dissected and carefully figured the so-called pro- 

 nephros of the Frog's tadpole, in a manner which as to accuracy 

 of detail anticipated later discovery. Again, in the early '8o's, 

 he had observed and recorded in a drawing the prae-pulmonary 

 aortic arch of the Amphibian, at a period antedating the re- 

 searches of Boas, which in connection with its discovery placed 

 the whole subject of the morphology of the pulmonary artery 

 of the vertebrata on its final basis, and brought harmony into 

 our ideas concerning it. 



Both these subjects lie at the root of modern advances in 

 vertebrate morphology. 



Concerning the skull, he was in the '8o's back to it with a 

 will. His line of attack was through the lampreys and hags and 

 the higher cartilaginous fishes, and he was following up a revo- 

 lutionary conception (already hinted at in his Hunterian Lec- 

 tures in 1864, and later in a Royal Society paper on Amphioxus 

 in 1875), that the trabeculae cranii, judged by their relation- 

 ships to the nerves, may represent a pair of prae-oral visceral 

 arches. In his unpublished notes there is evidence that he was 

 bringing to the support of this conclusion the discovery of a 

 supposed 4th branch to the trigeminal nerve — the relationships 

 of this (which he proposed to term the " hyporhinal " or palato- 

 nasal division) and the ophthalmic (to have been termed the 

 "orbitonasal"*) to the trabecular arch and a supposed pras- 



* A term already applied by him in 1875 to the corresponding nerve 

 in the Batrachia. (Ency. Brit, gth edition, vol. i., art. "Amphibia.") 



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