APPENDIX I 



475 



And well do I remember how, in the '8o's, both in the class- 

 room and in conversation, he would emphasise the fact that the 

 hypoglossus nerve roots of the mammal arise serially with the 

 ventral roots of the spinal nerves, little thinking that the dis- 

 covery by Froriep, in 1886, of their dorsal ganglionated counter- 

 parts, would establish the actual homology between the two, and 

 by leading to the conclusion that though actual vertebra; do 

 not contribute to the formation of the mammalian skull, its 

 occipital region is of truncal origin, mark the most revolution- 

 ary advance in cranial morphology since his own of 1856. 



Much of the final zoological work of his life lay with the 

 Bony Fishes, and he leaves unfinished (indeed only just com- 

 menced) a memoir embodying a new scheme of classification of 

 these, which shows that he was intending to do for them what 

 he did for Birds in the most active period of his career. It was 

 my good fortune to have helped as a hodman in the study of 

 these creatures, with a view to a Text-book we were to have 

 written conjointly, and as I realise what he was intending to 

 make out of the dry facts, I am filled with grief at the thought 

 of what we must have lost. His classification was based on the 

 labours of years, as testified by a vast accumulation of rough 

 notes and sketches, and as a conspicuous feature of it there 

 stands the embodiment under one head of all those fishes having 

 the swim-bladdei in connection with the auditory organ by 

 means of a chain of ossicles — a revolutionary arrangement, 

 which later, in the h^nds of the late Dr. Sagemahl, and by his 

 introduction of the famous term — " Ostariophyseae," has done 

 more than all else of recent years to clear the Ichthyological 

 air. Your father had anticipated this unpublished, and in a 

 proposal to unite the Herrings and Pikes into a single group, 

 the " Clupesoces," he had further given promise of a new sys- 

 tem, based on the study of the structure of the fins, jaws, and 

 reproductive organs of the Bony Fishes, the classifications of 

 which are still largely chaotic, which would have been as revo- 

 lutionary as it was rational. New terms both in taxonomy and 

 anatomy were contemplated, and in part framed. His published 

 terms " Elasmo-" and " Cysto-arian " are the adjective form of 

 t W o — far-reaching and significant — which give an idea of what 

 was to have come. Similarly, the spinose fin-rays were to have 

 been termed " acanthonemes," the branching and multiarticulate 



anything about the terms you mention. If you were to examine me 

 in my own papers I believe I should be plucked." 



