Teleutospores. 41 
surmounted by a prominent conical point. The membrane 
soon becomes dark and thickened, and each spore, en- 
veloped by its endospore, pretty well fills each compart- 
ment. In the mature state the entire teleutospore is 
clothed by a colourless transparent cuticle, which is gene- 
rally tuberculate. That this cuticle is distinct from the 
thick membrane is shown by warming a spore in caustic 
potash, when the former disappears, while the thick mem- 
brane remains unaffected. 
From two to four lateral germ canals or pores occur 
in the membrane, as Tulasne first showed (Plate IV. 
Fig. 5). These are arranged equatorially, and appear to 
be about four in number. Dietel,* however, remarks that 
in Ph. obtusum, Strauss, the germ-pores are placed as in 
Puccinia, namely, the superior one at the apex of the 
upper cell, and those of the lower cells laterally immediately 
below each of the septa, and that there is only one germ- 
pore in each cell. A similar condition, he states, exists in 
the Australian species Ph. barnardi, Plow. Longitudinal 
septation very rarely happens. ft 
In Coleosporium a similar condition exists, but in Tri- 
phragmium the divisions are multiple and longitudinal. 
In Cronartium the teleutospores are arranged in the form 
of a solid pillar or column, that projects perpendicularly 
to the spore-bed, and is surrounded at its base by a nest of 
uredospores. In Chrysomyxa the teleutospores occur in 
waxy masses, and consist of a series of superimposed 
spores, each of which has a single germ-pore placed as in 
the lower cell of Puccinia, not equatorially, but laterally 
near the upper end. In Gymnosporangium the spores are 
held together by a gelatinous matrix; they are shaped 
* Dietel, ‘* Beitrage zur’ Morphol. und Biolog. der Uredineen,” Botan. 
Centralblatt., bd. xxxii. (1887), tab. ii. figs. 1 and 2, reprint, p. 9. 
T Eysenhardt, Linnea,” vol. iii, 1828, 
