76 British Uvedinee and Ustilaginee. 
them passes into the other (Fig. 11). When the first pro- 
mycelial spore has become emptied of its contents, the 
second emits a tube which may remain as a germ-tube, or, 
as it becomes full of protoplasm, its end may swell out and 
form a third spore. (3) At a variable distance from the 
spore from which it arose a detached spore may form a 
connection with one of the segments of another promyce- 
lium ; sometimes as many as three spores may thus become 
united.* 
Germination, however, does not always occur in the 
above typical manner, namely, by the development of pro- 
mycelium and promycelial spores. From some of the 
largest teleutospores two promycelia are occasionally given 
off (Fig. 7). More commonly we find that, instead of pro- 
mycelial spores being produced in the regular manner 
above indicated, only one or two segments give rise to 
them. The others send off branches, into which their con- 
tents are emptied in the same manner as occurred when 
spores were formed. The free ends of the promycelial 
branches often come into contact with one another. When 
this happens they fuse together and become one continuous 
tube. (1) Thus a tube given off from one of the upper 
segments may form a connection with one of the lower 
segments of the same promycelium in the form of a bow 
(Fig. 4). (2) Two continuous segments may become united ° 
by forming what Brefeld calls a buckle-joint. This con- 
sists of the unequal growth of one side of the promycelium 
at the level of one of the septa; as this growing-out con- 
tinues the promycelium itself becomes bent at an angle, 
at first obtuse, but eventually acute. A reference to the 
figures (Plate VII. Figs. 3 and 9) will render this obvious. 
(3) Promycelia from two different spores may unite by 
branches in various ways, either by their ends or at any 
* Brefeld, loc. czt., pp. 54-67, t. ii, iii., figs, 1-17. 
