266 BRYOPHYTA 
styled an elaterophore. It is illustrated in the cases of JZefzgeria and 
Aneura (Fig. 127), and also in Pedlia (Fig. 128). In the former the 
elaterophore is attached internally at the distal end of the capsule, in the 
latter at its base. In Avewra the capsule-wall consists of two layers of 
cells, while the archesporium lies within. The differentiation of this body: 
is initiated early: in so young a sporogonium as that of Fig. 129 A 
there is a distinction already marked by the protoplasmic contents between 
a central group of more transparent cells and a peripheral band with 
j 
Aeoeg 0s 
SG Vane 
Fic. 127. Fic. 128. 
Aneura pinguts. Ripe capsule Lellia calycina. — Sporo- 
in longitudinal section. From the gonium opened, and emptied, 
summit the elaterophore hangs into showing the valves of the 
the spore-cavity, in which are many wall recurved, and an elatero- 
spores and_ elaters. Magnified. phore of many threads. (After 
(After Goebel.) Goebel.) 
denser protoplasm. The former constitutes only a central part of the 
elaterophore: it is clearly shown at a later stage that the differentiation 
has extended (Fig. 129 8), and that some of the inner products of the 
darker band shown in Fig. a are also developing as sterile cells, while 
it 1s only the fertile outer fringe which is the final residuum after these 
progressive steps of sterilisation. This point comes out even more clearly 
in a transverse section (Fig. 129 C), where the central group of cells 
first differentiated are readily distinguished from those differentiated later. 
There can be little doubt, after comparison with other Jungermanniaceae, 
that the history of the individual development in this case gives a correct 
