304 LYCOPODIALES 
similar in their general characters, as also in their clear definition, to 
those of the more differentiated types of Lycopodium or to Selagrnella. 
The general structure of the strobilus is, as in other Lycopods, essentially 
the same as that of the vegetative shoot, excepting in the presence of the 
sporangia. These are of very large size, and are commonly extended radially 
outwards from the axis, being interposed between the axis and the 
hgule: the latter then appears on the upper surface of the sporophyll, 
beyond the distal limit of the sporangium (Fig. 153). In Lepidodendron 
the cones thus constructed were borne on the ends of the ordinary 
Lepidophloios, sp. A =tangential section from the outside of a stem, passing through 
the leaf-bases, and showing their characteristic form, slightly enlarged. L=a single 
leaf-base, to show details; 4=collateral vascular bundle; Aa=the two parichnos- 
strands; @g=ligule in its pit. “10. Will. Coll., r9z7q4a. (After Scott.) 
branches; but in Szgz//arza they appear to have arisen laterally upon the 
main axis, from which after maturity they were deciduous, and each was 
borne upon an elongated pedicel covered with acicular bracts, while the 
cone itself showed a construction essentially similar to that of a small 
Lepidodendron. 
Though the type with a definite cone marked off from the sterile 
region was usual for the fossil Lycopodiales, it was not universal. In the 
imperfectly known plant, L/ewromora from the Trias, the whole main axis 
seems to have been a strobilus (Fig. 154), borne upon a Stigmarian base 
(compare Fig. 151). But a much more satisfactory example, from the 
Westphalian series (Middle Coal Measures) is that of Pinakodendron 
musivum, Weiss, specimens of which, discovered by M. Hector Delteure 
