MATONINEAE 565 
to hold a terminal position, has been. recognised as the inner branch of 
the second dichotomy. The segments themselves are pinnatifid, and the 
solitary sori are borne on their wings at points near to the midrib. The 
other species, AZ, sarmentosa, grows on rocks or on the branches of trees, 
with straggling, pendent leaves: at first sight the branching of the leaf 
seems quite different from that of JJ. pectinata; but this is due partly to 
the unequal development of the dichotomies, certain of the branches being 
represented only by arrested buds: partly it is due to their sympodial 
concatenation: but still the dichotomous branching appears to hold for 
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Fic. 315. 
Matonia pectinata, R. Br. Leaf drawn from a specimen in the British Museum 
Herbarium, by Mrs. Seward. } natural size. 
both. An interesting feature in this species is that the sori are borne in 
larger numbers, forming a row on either side of the distal part of the 
pinnule: an arrangement more closely resembling that in Gleichenia than 
that of Mf. pectinata. The structure of the sorus is, however, precisely 
like that in M. fectinata, and there is no doubt of the close alliance of 
the two species. 
SPORE-PRODUCING MEMBERS. 
The general structure of the mature sorus is well. known; the sporangia, 
commonly six to nine in number, form a simple ring-like series round the 
receptacle, and are covered till maturity by the thick and leathery hemi- 
spherical indusium, -which is ultimately deciduous. The orientation of 
the sporangia is not exactly uniform ; that of the majority is as in Glecchenza, 
but many have the annulus inclined, a consequence probably of crowding ; 
this is seen also in the fossil Zaccopteris. The annulus is incomplete at 
