34 AMPHISPORES 
AMPHISPORES. 
In countries which are arid or semi-arid, there is found in 
certain species a form of uredospore which has led to several 
mistakes owing to its misleading appearance. The spores are 
provided with a thick cell-wall or have the summit thickened 
conically, after the style of many a teleutospore of Uromyces, 
and are supported on a persistent pedicel, so that one would 
not take them for uredospores; nevertheless they will be found 
to have more than one germ-pore and to germinate by a germ- 
tube, although only after a period of rest. These were named 
by Carleton amphispores ; they were first discovered in Puccinia 
vexans. They are evidently a provision to enable the spore to 
pass through an unfavourable period unharmed, and reinfect a 
host of the same species when occasion arises. The amphi- 
spores of Puccinia atrofusca Holway, though echinulate and 
possessing two equatorial germ-pores, were first described by 
the discoverers, Dudley and Thompson, as the teleutospores of 
a Uromyces, and the same thing has happened in other cases, 
eg. Puccinia convolvulr, P. tosta, and P. cryptandri. The 
nearest approach to amphispores found in 
British species is in P. Pruni-spinosae (Fig. 
22), in which they have been mistaken for 
paraphyses and were so figured in a well- 
known text-book. There is another kind of 
spore, presently to be described, called a 
Hig, ee es mesospore, which bears a superficial resem- 
spinosae. x600.  blance to an amphispore. It isa mistake to 
call amphispores a transition-form between 
uredo- and teleutospores, since they are of later evolution than 
the two latter. 
TELEUTOSPORES. 
The meaning of the word teleutospore is end-spore ; it was 
considered to represent the stage when growth was ceasing for 
the season. This is not the case, however, in all species, and 
the word must now be used with another connotation, viz. a 
teleutospore is one which germinates by the production of a 
basidium and basidiospores. 
