126 UROMYCES 
Teleutospores, Sori generally hypophyllous, similar to the 
uredo-sori but more often confluent, 
always covered by the epidermis, 
! compact, shining, black; spores ovate- 
\ oblong, occasionally ellipsoid or pyri- 
form, rounded above, rarely truncate, 
often slightly thickened (up to 4,), 
Fig. 77. U.Dactylidis. Te. ‘Smooth, yellow-brown, darker only 
leutospores and the accom- glong the summit, 18—30 x 14— 
Eee eee 20; epispore 14m thick; pedicels 
brownish, persistent, nearly as long as the spore; paraphyses 
numerous, brown, agglutinated, dividing the sori into compart- 
ments. 
AKcidia on Ranunculus acris, R. bulbosus, R. repens, March— 
May; uredo- and teleutospores on Dactylis glomerata, from 
July onwards, often covering the leaves, less often the sheaths 
and culms, and persisting through the winter, especially on the 
latter. (Fig. 77.) 
This species and the following (U. Poae) are very closely allied, and 
should possibly be united. Perhaps more experimental cultures have been 
carried out with these two species than with most other Uredines; but the 
result is only a wild confusion of contradictory statements, from which 
one can infer, either that an immense number of intricately connected, but 
morphologically indistinguishable forms, inhabit the species of Ranun- 
cwlus and of Poa and Dactylis—or, preferably, that the factors which 
govern the success of an attempted infection are so numerous and so little 
known, that failure does not afford any ground for arriving at a definite 
conclusion. Those who wish to learn further should consult the long 
account of these results given in Sydow, Monographia, ii. pp. 312—16. 
In the British specimens of U. Dactylidis which I have examined, the 
paraphyses in the teleuto-sori, though often overlooked, are a conspicuous 
feature. But upon the question of paraphyses in the uredo-sori no agree- 
ment has been arrived at; the various authorities flatly contradict one 
another. Hither, therefore, the paraphyses occur differently in different 
countries, as Plowright suggests, or more than one species is included 
under this title, or their presence or absence is a matter of no importance. 
Against the latter suggestion, however, we must set the fact that in other 
cases, e.g. in Puccinia Sonchi, the paraphyses form a constant and dis- 
tinctive character. ; 
An ecidium occurring on Ranunculus acris belongs to Puceinia 
perplevans Plowr., but cannot be distinguished from the present one, 
