132 
PUCCINIA 
Uredospores. Sori generally hypophyllous, on irregular 
pallid-yellow or brownish spots, scattered or in 
clusters, about 1—14 mm. diam. often circi- 
nate, pulverulent, snuff-brown ; spores globose 
to ellipsoid, delicately echinulate, brown, 24— 
52 x 17—27 w, mostly with three germ-pores. 
[Teleutospores. Mixed with the uredospores, 
Fig.83. P. Chry- oblong or ellipsoid, rounded and_ slightly 
santhemi. Ure- 
dospore (Brit- 
ish). 
thickened above, usually rounded or some- 
h). what tapering at base, scarcely constricted, 
delicately verruculose, 
chestnut-brown, 35-—57 x 20—25 yz; 
pedicels thick, hyaline, persistent, 35—60 long; mesospores 
Fig. 84. P. Chrysanthemi. 
Teleutospores and uredo- 
spores, one abnormal (after 
Fischer). 
subglobose or pyriform, — slightly 
thickened at the summit, 32—387 x 
20-—21 p.] 
On leaves of Chrysanthemum in- 
dicum and C. sinense (not on other 
species of the genus, much less on 
other genera of Composite), in 
greenhouses, all the year round. The 
leaves that are attacked soon flag and 
die. (Figs. 88, 84.) 
This species is said to be very common 
in Japan. It was first observed in England 
in 1895, and has been found in other Euro- 
pean countries and in North America ; in 
1904 it reached Australia and New Zealand. 
In Japan it produces teleutospores in 
separate sori, which are hypophyllous, 
roundish, dark-brown and naked, but in 
Europe the teleutospores have been rarely 
seen, though mesospores occasionally occur. 
Abnormal and 2-celled uredospores (as well 
as 3- or 4-celled teleutospores) have been 
described and figured by Roze, Jacky and Fischer ; but these I have not 
seen in British specimens, 
Since, under the conditions in which the plants are grown here, the 
young shoots appear above ground before the old ones die away, it is 
probable that the parasite maintains itself by the uredospores alone; the 
alternative would be the possession of a perennial mycelium, which has not 
