194 PUCCINIA 
66. Puccinia Heraclei Grev. 
Trichobasis Heraclei Berk. ; Cooke, Micr. Fung. p. 225. 
Puccinia Heraclei Grev. Scot. Cr. Flor. pl. 42. Cooke, Handb. 
p. 502; Micr. Fung. p. 208. Sydow, Monogr. i. 387. Fischer, 
Ured. Schweiz, p. 132. 
P. Pimpinellae Strauss ; Plowr. Ured. p. 155 p.p. 
Spermogones. Amphigenous, scattered amongst the ecidia, 
pale-yellowish. 
4Aicidiospores. icidia hypophyllous, frequently on the 
petioles and especially on the nerves of the 
leaves, on thickened yellowish spots, densely 
crowded in irregular clusters, often causing 
distortion, between cup-shaped and pustulate, 
sometimes almost spherical and superficial ; 
peridium feebly developed, opening by a 
rounded pore; spores delicately verruculose, 
: yellowish, 21—32 x 18—28 p. 
ae gare ile Uredospores. Sori hi ttered 
clei. Aicidia on sp amphigenous, scattered, 
a (re- minute, chestnut-brown; spores globose to 
ellipsoid, densely echinulate, pale-brown, 25— 
32 x 19—27 yp, with three or four germ-pores. 
Teleutospores. Sori similar or more or less confluent on the 
nerves, surrounded by the ferruginous epidermis, pulverulent, 
blackish ; spores ellipsoid, rounded at both ends, hardly con- 
stricted, reticulated, brown, 26—387 x 18—27 w; pedicels hya- 
line, short, deciduous. 
On Heracleum Sphondyliwm. Aicidia, March—June; teleu- 
tospores, August. Not common. (Fig. 142.) 
This species closely resembles P. Chaerophylli, but is distinguished by 
its less densely reticulated teleutospores. Semadeni proved by experi- 
ment that they are distinct species, but no one has as yet reared all the 
spore-forms of P. Heraclei from the basidiospores, as has been done 
for P. Chaerophylli. The ecidia of this species are more conspicuous 
than those of its allies; they occur in swollen patches, reminding one 
of the acidia on Smyrnium Olusatrum, and being sometimes almost 
spherical and superficial might be compared to a group of miniature Peziza 
vesiculosa. 
DISTRIBUTION: Central and North-western Europe. 
