ON POLYGONACE 225 
95. Puccinia Conopodii-Bistorte Kleb. 
Puccinia Bistortae DC. Flor. fr. vi. 61. Cooke, Micr. Fung. p. 204; 
Grevillea, ii. 161. Plowr. Ured. p. 192 (7). Sydow, Monogr. 
i. 571 (all pro parte). Soppitt, Grevillea, xxii. 45 ; Gard. Chron. 
1895, xviii. 773. 
P. Conopodii- Bistortae Kleb. Wirtsw. Rostp. p. 318. 
Acidiospores. AKcidia on large swellings of a bright orange 
colour, immersed, spherical or flat, not at all cup-shaped, margin 
not projecting; spores delicately verruculose, orange, 15—20 y. 
Uredospores. Sori hypophyllous, minute, roundish, yellowish- 
red, soon naked; spores globose to shortly ellipsoid, finely 
echinulate, pale yellowish-brown, 21—24 w. 
Teleutospores. Sori hypophyllous, scattered or united in 
roundish groups, soon naked and pulverulent, dark- 
brown; spores oblong or subclavate, rounded at ae 
both ends or slightly oblique at the apex, faintly ( 
constricted, clear yellowish-brown, 28—42 x 16— ey) 
25; epispore equally thick, smooth, but some- 
times marked with a very few longitudinal or 5, 178. 
oblique rows of delicate warts; no papilla. on  P. Conopodii- 
the germ-pores; pedicels hyaline, short, very tee 
deciduous. 
ficidia on Conopodiuwm denudatum, Yorkshire, May; teleuto- 
spores on Polygonum Bistorta, June—August, not common. 
(Fig. 173.) 
The Puccinias on Pol. Bistorta are not yet well known. There appear to: 
be at least four (or five) distinct forms, which are divisible morphologically 
into two groups—(1) those whose teleutospores have no apical papilla, 
(2) those which have a small hemispherical papilla at the summit; the 
latter are called by Sydow in the Monographia P. mammillata Schrét. 
(Pilz. Schles. p. 340). They are all hetercecious. 
Of the former group, one form has an ecidium on Conopodium, and its 
other spore-stages on P. Bistorta. Itis the one mentioned above ; the life: 
history was first demonstrated by Soppitt (2.¢.). It is remarkable as 
being the first instance known of a hetercecious Puccinia that had its 
teleutospores on a Dicotyledon. The second form has its ecidium on 
Angelica silvestris and Carum Caruz and its teleutospores on Pol. Bistorta 
and Pol. viviparum. It would perhaps be better divided into P. Angelicae- 
Bistortae Web. (= P. Cari-Bistortae) and P. Polygoni-vivipari Karst. The 
connection in the former has been demonstrated by Klebahn and Fischer ; 
G.U. 15 
