260 PUCCINIA 
122. Puccinia dispersa (sens. lat.) Er. et Henn. 
Trichobasis Rubigo-vera Lév.; Cooke, Micr. Fung. p. 222, pl. 7, 
f. 140—2 (?). 
Puccinia straminis Cooke, Mier. Fung. p. 202 p.p. 
P. Rubigo-vera Plowr. Ured. p. 167 p.p. Sace. Syll. vii. 624 p.p. 
P. dispersa Erikss. et Henning, Getreideroste, p. 210 (1896). Trans. 
Brit. Myc. Soe. i. 58. 
Uredospores. Sori generally epiphyllous or a few fhypo- 
phyllous, 1—2 mm. long, scattered without order, rarely con- 
fluent, oblong or punctiform, rust-coloured or dirty-ochre, be- 
coming paler; spores more or less globose, shortly echinulate, 
dirty-yellow or dull-orange, 16—28; membrane distinctly 
brownish (pale chocolate-umber) when mature; germ-pores 
7—10, scattered over the whole surface. 
Teleutospores. Sori hypophyllous or less often on the 
sheaths, scattered or slightly and irregularly aggregated, rarely 
in distinct lines, small, oblong, covered by the epidermis, black ; 
spores oblong to clavate, truncate, rounded, or obtusely and 
obliquely pointed above, slightly thickened, gently constricted, 
narrowed downwards, smooth, brown, 35—56 x 12—23 uw; pedi- 
cels short ; paraphyses numerous, brownish, more or less curved, 
surrounding the spores. 
This is a general description of the forms included under the name 
Brown Rust, to which the title P. dispersa was originally given. The 
dirty-orange colour of the uredospores, which distinguishes them at a 
glance from P. glumarum, is due to the fact that the membrane of the 
spores is brownish, not hyaline ; the spore contents are orange in colour. 
The germ-pores are scarcely perceptible in the immature or untreated 
spore, but they can be seen easily if a spore is squeezed strongly between 
the cover-glass and the slide, or by choosing a mature and empty spore. 
In the teleutospores only the upper slightly thickened wall is dark- 
chestnut, the rest being thin-walled and pale; there is usually also a 
chestnut-brown band at the apex of the pedicel. The structures called 
paraphyses here in the teleuto-sori are quite different from those called by 
the samehame in Melampsora, etc. ; they are erect, coherent, thick-walled, 
prismatic cells, which surround the teleuto-sori, or in the case of the 
larger ones divide them into loculi. There are also paraphyses of the 
ordinary shape, with a brownish membrane, mingled with the uredospores 
in certain cases, but the occurrence of these seems, so far as is known at 
present, to be somewhat fortuitous, 
